Drupe
Encyclopedia
Botany
Botany, plant science, or plant biology is a branch of biology that involves the scientific study of plant life. Traditionally, botany also included the study of fungi, algae and viruses...
, a drupe is a fruit
Fruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...
in which an outer fleshy part (exocarp, or skin; and mesocarp, or flesh) surrounds a shell (the pit, stone or pyrene
Pyrena
Pyrena or pyrene is the name for a nutlet resembling a seed, or the kernel of a drupe or drupelet....
) of hardened endocarp with a seed
Seed
A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...
inside. These fruits develop from a single carpel, and mostly from flower
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...
s with superior ovaries. The definitive characteristic of a drupe is that the hard, lignified stone (or pit) is derived from the ovary wall of the flower. In an aggregate fruit composed of small, individual drupes, each individual is termed a drupelet.
Other fleshy fruits may have a stony enclosure that comes from the seed coat surrounding the seed, but such fruits are not drupes.
Some flowering plant
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...
s that produce drupes are coffee
Coffea arabica
Coffea arabica is a species of Coffea originally indigenous to the mountains of Yemen in the Arabian Peninsula, hence its name, and also from the southwestern highlands of Ethiopia and southeastern Sudan. It is also known as the "coffee shrub of Arabia", "mountain coffee" or "arabica coffee"...
, jujube
Jujube
Ziziphus zizyphus , commonly called jujube , red date, Chinese date, Korean date, or Indian date is a species of Ziziphus in the buckthorn family Rhamnaceae, used primarily as a fruiting shade tree.-Distribution:Its precise natural distribution is uncertain due to extensive cultivation,...
, mango
Mango
The mango is a fleshy stone fruit belonging to the genus Mangifera, consisting of numerous tropical fruiting trees in the flowering plant family Anacardiaceae. The mango is native to India from where it spread all over the world. It is also the most cultivated fruit of the tropical world. While...
, olive
Olive
The olive , Olea europaea), is a species of a small tree in the family Oleaceae, native to the coastal areas of the eastern Mediterranean Basin as well as northern Iran at the south end of the Caspian Sea.Its fruit, also called the olive, is of major agricultural importance in the...
, most palms (including date
Date Palm
The date palm is a palm in the genus Phoenix, cultivated for its edible sweet fruit. Although its place of origin is unknown because of long cultivation, it probably originated from lands around the Persian Gulf. It is a medium-sized plant, 15–25 m tall, growing singly or forming a clump with...
, coconut
Coconut
The coconut palm, Cocos nucifera, is a member of the family Arecaceae . It is the only accepted species in the genus Cocos. The term coconut can refer to the entire coconut palm, the seed, or the fruit, which is not a botanical nut. The spelling cocoanut is an old-fashioned form of the word...
and oil palm
Oil palm
The oil palms comprise two species of the Arecaceae, or palm family. They are used in commercial agriculture in the production of palm oil. The African Oil Palm Elaeis guineensis is native to West Africa, occurring between Angola and Gambia, while the American Oil Palm Elaeis oleifera is native to...
s), pistachio
Pistachio
The pistachio, Pistacia vera in the Anacardiaceae family, is a small tree originally from Persia , which now can also be found in regions of Syria, Lebanon, Turkey, Greece, Kyrgyzstan, Turkmenistan, India, Pakistan, Egypt, Sicily and possibly Afghanistan , as well as in the United States,...
, white sapote
White sapote
White sapote, Casimiroa edulis, also known as cochitzapotl in the Nahuatl language is a species of tropical fruiting tree in the family Rutaceae, native to eastern Mexico and Central America south to Costa Rica....
, and all members of the genus Prunus
Prunus
Prunus is a genus of trees and shrubs, which includes the plums, cherries, peaches, apricots and almonds. There are around 430 species spread throughout the northern temperate regions of the globe. Many members of the genus are widely cultivated for fruit and ornament.-Botany:Members of the genus...
, including the almond
Almond
The almond , is a species of tree native to the Middle East and South Asia. Almond is also the name of the edible and widely cultivated seed of this tree...
(in which the mesocarp is somewhat leathery). apricot
Apricot
The apricot, Prunus armeniaca, is a species of Prunus, classified with the plum in the subgenus Prunus. The native range is somewhat uncertain due to its extensive prehistoric cultivation.- Description :...
, cherry
Cherry
The cherry is the fruit of many plants of the genus Prunus, and is a fleshy stone fruit. The cherry fruits of commerce are usually obtained from a limited number of species, including especially cultivars of the wild cherry, Prunus avium....
, damson
Damson
The damson or damson plum is an edible drupaceous fruit, a subspecies of the plum tree. Sometimes called the Damask plum, damsons are commonly used in the preparation of jams and jellies...
, nectarine, peach
Peach
The peach tree is a deciduous tree growing to tall and 6 in. in diameter, belonging to the subfamily Prunoideae of the family Rosaceae. It bears an edible juicy fruit called a peach...
, and plum
Plum
A plum or gage is a stone fruit tree in the genus Prunus, subgenus Prunus. The subgenus is distinguished from other subgenera in the shoots having a terminal bud and solitary side buds , the flowers in groups of one to five together on short stems, and the fruit having a groove running down one...
.
Corking is a nutritional disorder in stone fruit caused by a lack of boron
Boron
Boron is the chemical element with atomic number 5 and the chemical symbol B. Boron is a metalloid. Because boron is not produced by stellar nucleosynthesis, it is a low-abundance element in both the solar system and the Earth's crust. However, boron is concentrated on Earth by the...
and/or calcium
Calcium
Calcium is the chemical element with the symbol Ca and atomic number 20. It has an atomic mass of 40.078 amu. Calcium is a soft gray alkaline earth metal, and is the fifth-most-abundant element by mass in the Earth's crust...
.
Terminology
The term stone fruit (also stonefruit) can be a synonym for drupe or, more typically, it can mean just the fruit of the PrunusPrunus
Prunus is a genus of trees and shrubs, which includes the plums, cherries, peaches, apricots and almonds. There are around 430 species spread throughout the northern temperate regions of the globe. Many members of the genus are widely cultivated for fruit and ornament.-Botany:Members of the genus...
genus.
Freestone refers to a drupe having a stone which can be removed from the flesh with ease. The flesh is not attached to the stone and does not need to be cut to free the stone. Freestone varieties of fruits are preferred for uses that require careful removal of the stone, especially if removal will be done by hand. Freestone plums are preferred for making homegrown prune
Prune
A prune is any of various plum cultivars, mostly Prunus domestica or European Plum, sold as fresh or dried fruit. The dried fruit is also referred to as a dried plum...
s, and freestone sour cherries
Sour Cherry
Prunus cerasus, or the sour cherry, is a species of Prunus in the subgenus Cerasus , native to much of Europe and southwest Asia. It is closely related to the wild cherry Prunus cerasus, or the sour cherry, is a species of Prunus in the subgenus Cerasus (cherries), native to much of Europe and...
are preferred for making pies and cherry soup.
Clingstone refers to a drupe having a stone which cannot easily be removed from the flesh. The flesh is attached strongly to the stone and must be cut to free the stone. Clingstone varieties of fruits in the genus Prunus are preferred as table fruit and for jams, because the flesh of clingstone fruits tends to be more tender and juicy throughout.
Tryma is a specialized term for such nut
Nut (fruit)
A nut is a hard-shelled fruit of some plants having an indehiscent seed. While a wide variety of dried seeds and fruits are called nuts in English, only a certain number of them are considered by biologists to be true nuts...
-like drupes that are difficult to categorize. Hickory nuts (Carya) and walnuts (Juglans) in the Juglandaceae
Juglandaceae
The Juglandaceae, also known as the Walnut Family, is a family of trees, or sometimes shrubs, in the order Fagales. Various members of this family are native to the Americas, Eurasia, and Southeast Asia....
family grow within an outer husk; these fruits are technically drupes or drupaceous nuts, and thus not true botanical nuts.
Ecology
Drupes, with their sweet, fleshy outer layer, attract the attention of animals as a foodFood
Food is any substance consumed to provide nutritional support for the body. It is usually of plant or animal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals...
, and the plant population benefits from the resulting dispersal of its seeds
Seed dispersal
Seed dispersal is the movement or transport of seeds away from the parent plant. Plants have limited mobility and consequently rely upon a variety of dispersal vectors to transport their propagules, including both abiotic and biotic vectors. Seeds can be dispersed away from the parent plant...
. The endocarp (pit or stone) is sometimes dropped after the fleshy part is eaten, but is often swallowed, passing through the digestive tract, and returned to the soil in feces
Feces
Feces, faeces, or fæces is a waste product from an animal's digestive tract expelled through the anus or cloaca during defecation.-Etymology:...
with the seed inside unharmed. This passage through the digestive tract can reduce the thickness of the endocarp, and thus can aid in germination rates. The process is known as scarification
Scarification (botany)
Scarification in botany involves cutting the seed coat using abrasion, thermal stress, or chemicals to encourage germination.-In botany:Scarification is a natural process important for germination of many species' seeds. This process involves the breaching of the natural seed coating by mechanical,...
.
Examples
Typical drupes include peaches, plums, and cherries (see pictures below).The coconut
Coconut
The coconut palm, Cocos nucifera, is a member of the family Arecaceae . It is the only accepted species in the genus Cocos. The term coconut can refer to the entire coconut palm, the seed, or the fruit, which is not a botanical nut. The spelling cocoanut is an old-fashioned form of the word...
is also a drupe, but the mesocarp is fibrous or dry (termed a husk), so this type of fruit is classified as a simple dry fruit, fibrous drupe. Unlike other drupes, the coconut seed is unlikely to be dispersed by being swallowed by fauna, due to its large size. It can, however, float extremely long distances across oceans.
Bramble
Bramble
Brambles are thorny plants of the genus Rubus, in the rose family . Bramble fruit is the fruit of any such plant, including the blackberry and raspberry. The word comes from Germanic *bram-bezi, whence also German Brombeere , Dutch Braam and French framboise...
fruits (such as the blackberry
Blackberry
The blackberry is an edible fruit produced by any of several species in the Rubus genus of the Rosaceae family. The fruit is not a true berry; botanically it is termed an aggregate fruit, composed of small drupelets. The plants typically have biennial canes and perennial roots. Blackberries and...
or the raspberry
Raspberry
The raspberry or hindberry is the edible fruit of a multitude of plant species in the genus Rubus, most of which are in the subgenus Idaeobatus; the name also applies to these plants themselves...
) are aggregates of drupelets. The fruit of blackberries and raspberries comes from a single flower whose pistil is made up of a number of free carpels. However, mulberries
Mulberry
Morus is a genus of flowering plants in the family Moraceae. The 10–16 species of deciduous trees it contains are commonly known as Mulberries....
, which closely resemble blackberries, are not aggregate fruit, but are multiple fruit
Multiple fruit
Multiple fruits are fruits that are formed from a cluster of flowers . Each flower in the inflorescence produces a fruit, but these mature into a single mass...
s, actually derived from bunches of catkin
Catkin
A catkin or ament is a slim, cylindrical flower cluster, with inconspicuous or no petals, usually wind-pollinated but sometimes insect pollinated . They contain many, usually unisexual flowers, arranged closely along a central stem which is often drooping...
s, each drupelet thus belonging to a different flower.
Certain drupes occur in large clusters, as in the case of palm species, where a sizable array of drupes are found in a cluster. Examples of such large drupe clusters include Jubaea chilensis in central Chile
Chile
Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...
and Washingtonia filifera
Washingtonia filifera
Washingtonia filifera , with the common names California Fan Palm , Desert Fan Palm, Cotton palm, and Arizona Fan Palm. It is a palm native to southwestern North America between an elevation range of , at seeps, desert bajadas, and springs where underground water is continuously available...
in the Sonoran Desert
Sonoran Desert
The Sonoran Desert is a North American desert which straddles part of the United States-Mexico border and covers large parts of the U.S. states of Arizona and California and the northwest Mexican states of Sonora, Baja California, and Baja California Sur. It is one of the largest and hottest...
of North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...
.