Liaoningopterus
Encyclopedia
Liaoningopterus, sometimes misspelled "Liaoningopteryx", was a genus
of ornithocheirid
pterodactyloid pterosaur
from the Barremian
-Aptian
-age Lower Cretaceous Jiufotang Formation
of Chaoyang, Liaoning
, China
.
The genus was named in 2003 by Wang Xiaolin and Zhou Zhimin. The type species
is Liaoningopterus gui. The genus name is derived from Liaoning
and a Latinised Greek pteron, "wing". The specific name honours Professor Gu Zhiwei, an invertebrate
specialist who has pioneered the study of the Jehol Biota
.
The genus is based on holotype
IVPP V-13291
, a crushed partial skull and skeleton including the jaws, teeth, a cervical
vertebra, and bones of the finger
supporting the wing. It was a large pterosaur — the largest known from China at the time of description — with a skull length estimated at 61 cm (24 in), and a wingspan
estimated at five metres (16.4 ft). The skull was long and low, bearing low crests close to the tip of the beak on both lower and upper jaws. The snout crest was twelve centimetres long, was symmetrical in form and had a maximum height of seventeen millimetres. The edge of the upper jaw was very straight. The teeth were only found at the anterior end of the jaws. They were elongated but robust, generally increasing in size from the back to the front. The fourth tooth in the upper jaw is with a length of 81 millimetres the largest known for any pterosaur. It is exceptional in size compared to the other teeth of Liaoningopterus also, the longest tooth in the lower jaw having a length of 41 millimetres. Tooth length in the specimen is very variable, which the authors explained by the presence of recently erupted replacement teeth. There were twenty pairs of teeth in the upper jaws and thirteen or fourteen pairs in the lower jaws.
The preserved cervical vertebra has a centrum length of 46 millimetres and a centrum height of 34 millimetres. From the wing bones pieces of the first phalanx can be recognised which had an estimated total length of about fifty centimetres.
The authors described Liaoningopterus as being probably a piscivore
, due to the long, pointed snout.
Wang classified Liaoningopterus as a member of the Anhangueridae, mainly because of the crests. This opinion was restated by him in 2005. In 2006 Lü Junchang published a cladistic analysis showing Liaoningopterus to be a basal member of the Anhangueridae; in 2008 an analysis by Ji Qiang had Liaoningopterus in a trichotomy
with Anhanguera
and Tropeognathus. Within the alternative terminology of David Unwin Liaoningopterus is a member of the Ornithocheiridae
.
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...
of ornithocheirid
Ornithocheiridae
Ornithocheiridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth.-Classification:Listing of genera after Unwin , except where noted.* Family Ornithocheiridae** ?Aetodactylus...
pterodactyloid pterosaur
Pterosaur
Pterosaurs were flying reptiles of the clade or order Pterosauria. They existed from the late Triassic to the end of the Cretaceous Period . Pterosaurs are the earliest vertebrates known to have evolved powered flight...
from the Barremian
Barremian
The Barremian is an age in the geologic timescale between 130.0 ± 1.5 Ma and 125.0 ± 1.0 Ma). It is a subdivision of the Early Cretaceous epoch...
-Aptian
Aptian
The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous epoch or series and encompasses the time from 125.0 ± 1.0 Ma to 112.0 ± 1.0 Ma , approximately...
-age Lower Cretaceous Jiufotang Formation
Yixian Formation
The Yixian Formation is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans 11 million years during the early Cretaceous period...
of Chaoyang, Liaoning
Liaoning
' is a province of the People's Republic of China, located in the northeast of the country. Its one-character abbreviation is "辽" , a name taken from the Liao River that flows through the province. "Níng" means "peace"...
, China
People's Republic of China
China , officially the People's Republic of China , is the most populous country in the world, with over 1.3 billion citizens. Located in East Asia, the country covers approximately 9.6 million square kilometres...
.
The genus was named in 2003 by Wang Xiaolin and Zhou Zhimin. The type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...
is Liaoningopterus gui. The genus name is derived from Liaoning
Liaoning
' is a province of the People's Republic of China, located in the northeast of the country. Its one-character abbreviation is "辽" , a name taken from the Liao River that flows through the province. "Níng" means "peace"...
and a Latinised Greek pteron, "wing". The specific name honours Professor Gu Zhiwei, an invertebrate
Invertebrate
An invertebrate is an animal without a backbone. The group includes 97% of all animal species – all animals except those in the chordate subphylum Vertebrata .Invertebrates form a paraphyletic group...
specialist who has pioneered the study of the Jehol Biota
Jehol Biota
The Jehol Biota includes all the living organisms - the ecosystem - of northeastern China between 133 to 120 million years ago. This is the Lower Cretaceous ecosystem which left fossils in the Yixian Formation and Jiufotang Formation. It is also believed to have left fossils in the Sinuiju series...
.
The genus is based on holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...
IVPP V-13291
Institute of Vertebrate Paleontology and Paleoanthropology
The Institute of Vertebrate Paleontology and Paleoanthropology of China is a prominent research institution and collections repository for fossils, including many dinosaur and pterosaurand cat poo specimens...
, a crushed partial skull and skeleton including the jaws, teeth, a cervical
Neck
The neck is the part of the body, on many terrestrial or secondarily aquatic vertebrates, that distinguishes the head from the torso or trunk. The adjective signifying "of the neck" is cervical .-Boner anatomy: The cervical spine:The cervical portion of the human spine comprises seven boney...
vertebra, and bones of the finger
Finger
A finger is a limb of the human body and a type of digit, an organ of manipulation and sensation found in the hands of humans and other primates....
supporting the wing. It was a large pterosaur — the largest known from China at the time of description — with a skull length estimated at 61 cm (24 in), and a wingspan
Wingspan
The wingspan of an airplane or a bird, is the distance from one wingtip to the other wingtip. For example, the Boeing 777 has a wingspan of about ; and a Wandering Albatross caught in 1965 had a wingspan of , the official record for a living bird.The term wingspan, more technically extent, is...
estimated at five metres (16.4 ft). The skull was long and low, bearing low crests close to the tip of the beak on both lower and upper jaws. The snout crest was twelve centimetres long, was symmetrical in form and had a maximum height of seventeen millimetres. The edge of the upper jaw was very straight. The teeth were only found at the anterior end of the jaws. They were elongated but robust, generally increasing in size from the back to the front. The fourth tooth in the upper jaw is with a length of 81 millimetres the largest known for any pterosaur. It is exceptional in size compared to the other teeth of Liaoningopterus also, the longest tooth in the lower jaw having a length of 41 millimetres. Tooth length in the specimen is very variable, which the authors explained by the presence of recently erupted replacement teeth. There were twenty pairs of teeth in the upper jaws and thirteen or fourteen pairs in the lower jaws.
The preserved cervical vertebra has a centrum length of 46 millimetres and a centrum height of 34 millimetres. From the wing bones pieces of the first phalanx can be recognised which had an estimated total length of about fifty centimetres.
The authors described Liaoningopterus as being probably a piscivore
Piscivore
A piscivore is a carnivorous animal which eats primarily fish. Piscivory was the diet of early tetrapods , insectivory came next, then in time reptiles added herbivory....
, due to the long, pointed snout.
Wang classified Liaoningopterus as a member of the Anhangueridae, mainly because of the crests. This opinion was restated by him in 2005. In 2006 Lü Junchang published a cladistic analysis showing Liaoningopterus to be a basal member of the Anhangueridae; in 2008 an analysis by Ji Qiang had Liaoningopterus in a trichotomy
Trichotomy
In mathematics, the Law of Trichotomy states that every real number is either positive, negative, or zero. More generally, trichotomy is the property of an order relation...
with Anhanguera
Anhangüera
Anhangüera originates from a word in the Tupi language which was the nickname given by Brazilian Indians to an explorer and "bandeirante", Bartolomeu Bueno da Silva .It may also refer to:* Anhanguera, Goiás, a municipality in the state of Goiás...
and Tropeognathus. Within the alternative terminology of David Unwin Liaoningopterus is a member of the Ornithocheiridae
Ornithocheiridae
Ornithocheiridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth.-Classification:Listing of genera after Unwin , except where noted.* Family Ornithocheiridae** ?Aetodactylus...
.
External links
- Liaoningopterus in The Pterosauria