List of caneberries diseases
Encyclopedia
Bacterial diseases
Bacterial diseases |
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Crown and cane gall | Agrobacterium tumefaciens Agrobacterium tumefaciens Agrobacterium tumefaciens is the causal agent of crown gall disease in over 140 species of dicot. It is a rod shaped, Gram negative soil bacterium... Agrobacterium rubi |
Fire blight | Erwinia amylovora |
Hairy root | Agrobacterium rhizogenes Agrobacterium rhizogenes Agrobacterium rhizogenes is a Gram negative soil bacterium that produces hairy root disease in dicotyledonous plants.A... |
Leafy gall | Rhodococcus fascians Rhodococcus fascians Rhodococcus fascians is a Gram positive bacterial phytopathogen that causes leafy gall disease. R. fascians is the only phytopathogenic member of the Rhodococcus genus; its host range includes both dicotyledonous and monocotyledonous hosts... = Corynebacterium fascians |
Pseudomonas blight | Pseudomonas syringae Pseudomonas syringae Pseudomonas syringae is a rod shaped, Gram-negative bacterium with polar flagella. It is a plant pathogen which can infect a wide range of plant species, and exists as over 50 different pathovars, all of which are available to legitimate researches via international culture collections such as the... |
Fungal diseases
Fungal diseases |
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Anthracnose | Elsinoë veneta Elsinoë veneta Elsinoë veneta is a plant pathogen, who is the causal agent of anthracnose of raspberry. -External links:* *... Sphaceloma necator [anamorph] |
Armillaria root rot Shoestring root rot |
Armillaria mellea Armillaria mellea Armillaria mellea is a plant pathogen and a species of Honey fungus. It causes Armillaria root rot in many plant species. The mushrooms are edible but some people may be intolerant to them. The fungus produces mushrooms around the base of trees it has infected... Rhizomorpha subcorticalis Rhizomorpha subcorticalis Rhizomorpha subcorticalis is a fungus that is a plant pathogen that causes root rot in several species, including strawberries and avocado.- External links :* *... [anamorph] |
Ascospora dieback | Clethridium corticola Ascospora ruborum Ascospora ruborum Ascospora ruborum is an ascomycete fungus that is a plant pathogen that causes dieback on Rubus sp..- External links :* *... Seimatosporium lichenicola [anamorph] |
Blackberry rust | Phragmidium violaceum Phragmidium violaceum Phragmidium violaceum is a plant pathogen native to Europe, Africa, and the Middle East. It primarily infects Rubus species.It has been used in the biological control of invasive blackberry species in Chile, Australia, and New Zealand. In 2005, it was discovered growing on Himilayan blackberry... |
Black rot | Phyllosticta carpogena Phyllosticta carpogena Phyllosticta carpogena is a plant pathogen.- External links :**... |
Blotch | Mycosphaerella confusa Mycosphaerella confusa Mycosphaerella confusa is a plant pathogen.-External links:* *... Pseudocercospora rubi [anamorph] Rhabdospora ramealis |
Botryosphaeria cane canker (blackberry) | Botryosphaeria dothidea Botryosphaeria dothidea Botryosphaeria dothidea is a plant pathogen that causes the formation of cankers on a wide variety of tree species.- External links :**... Fusicoccum aesculi Fusicoccum aesculi Fusicoccum aesculi is a fungus. It is a plant pathogen.- External links :**... [anamorph] |
Botrytis fruit rot and blossom blight = gray mold | Botrytis cinerea Botrytis cinerea Botrytis cinerea is a necrotrophic fungus that affects many plant species, although its most notable hosts may be wine grapes. In viticulture, it is commonly known as botrytis bunch rot; in horticulture, it is usually called grey mould or gray mold.The fungus gives rise to two different kinds of... Botryotinia fuckeliana Botryotinia fuckeliana Botryotinia fuckeliana is a plant pathogen, and the causal agent of gray mold disease.The specific epithet was named by mycologist Heinrich Anton de Bary in honor of another mycologist, Karl Wilhelm Gottlieb Leopold Fuckel.... [teleomorph] |
Boysenberry decline | Cercosporella rubi Cercosporella rubi Cercosporella rubi is a plant pathogen which causes rossette disease of blackberry. The disease is a particular problem in the southeast of the USA. The disease causes witch's broom symptoms as well as double blossoms, and can severely reduce yield.... |
Cane and leaf rust | Kuehneola uredinis Kuehneola uredinis Kuehneola uredinis is a plant pathogen.... |
Cane blight | Diapleella coniothyrium Leptosphaeria coniothyrium Leptosphaeria coniothyrium Leptosphaeria coniothyrium is a plant pathogen. It can be found across the world. - External links :* *... Coniothyrium fuckelii [anamorph] |
Cane Botrytis | Botrytis cinerea Botrytis cinerea Botrytis cinerea is a necrotrophic fungus that affects many plant species, although its most notable hosts may be wine grapes. In viticulture, it is commonly known as botrytis bunch rot; in horticulture, it is usually called grey mould or gray mold.The fungus gives rise to two different kinds of... Botryotinia fuckeliana Botryotinia fuckeliana Botryotinia fuckeliana is a plant pathogen, and the causal agent of gray mold disease.The specific epithet was named by mycologist Heinrich Anton de Bary in honor of another mycologist, Karl Wilhelm Gottlieb Leopold Fuckel.... [teleomorph] |
Downy mildew = dryberry | Peronospora sparsa Peronospora sparsa Peronospora sparsa is a plant pathogen that causes downy mildew on rose.- External links :**... = Peronospora rubi |
Dryberry disease (loganberry) | Rhizoctonia rubi Rhizoctonia rubi Rhizoctonia rubi is a fungus that is a plant pathogen.-External links:**... (plus dryberry mite, see under miscellaneous disorders) |
Fruit rots, minor | Alternaria Alternaria Alternaria is a genus of ascomycete fungi. Alternaria species are known as major plant pathogens. They are also common allergens in humans, growing indoors and causing hay fever or hypersensitivity reactions that sometimes lead to asthma... spp. Cladosporium Cladosporium Cladosporium is a genus of fungi including some of the most common indoor and outdoor molds. Species produce olive-green to brown or black colonies, and have dark-pigmented conidia that are formed in simple or branching chains.... spp. Colletotrichum gloeosporioides Colletotrichum acutatum Colletotrichum acutatum Colletotrichum acutatum is a plant pathogen.- External links :* *... Penicillium Penicillium Penicillium is a genus of ascomycetous fungi of major importance in the natural environment as well as food and drug production. Members of the genus produce penicillin, a molecule that is used as an antibiotic, which kills or stops the growth of certain kinds of bacteria inside the body... spp. |
Gnomonia cane canker | Gnomonia rubi Gnomonia rubi Gnomonia rubi is a plant pathogen that causes cane canker on Rubus.- External links :... |
Late leaf rust | Pucciniastrum americanum Pucciniastrum americanum Pucciniastrum americanum is a plant pathogen.- External links :**... Pucciniastrum arcticum Pucciniastrum arcticum Pucciniastrum arcticum is a plant pathogen.- External links :**... |
Leaf spots | Discohainesia oenotherae Hainesia lythri [anamorph] Mycosphaerella Mycosphaerella Mycosphaerella is a genus of sac fungi. With more than 10,000 species, it is the largest genus of plant pathogen fungi.... spp. Phyllosticta Phyllosticta Phyllosticta is a genus of fungi. Many of the species in this genus are plant pathogens.-Species:*Phyllosticta aberiae*Phyllosticta abietis*Phyllosticta abortiva*Phyllosticta abramovii*Phyllosticta abricola... spp. |
Nectria canker (raspberry) | Nectria mammoidea var. rubi Nectria mammoidea var. rubi Nectria mammoidea var. rubi is a plant pathogen.-External links:* *... Cylindrocarpon ianthothele var. ianthothele Cylindrocarpon ianthothele var. ianthothele Cylindrocarpon ianthothele var. ianthothele is a fungal plant pathogen.-External links:* *... [anamorph] |
Orange rust | Arthuriomyces peckianus Arthuriomyces peckianus Arthuriomyces peckianus is a fungal plant pathogen, which causes orange rust on members of the genus Rubus, and various species of berries. It is found in central and eastern North America, and Eurasia.-External links:*... (long-cycled rust) Gymnoconia nitens Gymnoconia nitens Gymnoconia nitens is a species of rust fungus in the Phragmidiaceae family. It is a plant pathogen, and causes orange rust on various berries. The species was originally described in 1822 by mycologist Lewis David de Schweinitz as Aecidium luminatum.... (short-cycled rust) |
Phytophthora root rot | Phytophthora Phytophthora Phytophthora is a genus of plant-damaging Oomycetes , whose member species are capable of causing enormous economic losses on crops worldwide, as well as environmental damage in natural ecosystems. The genus was first described by Heinrich Anton de Bary in 1875... spp. Phytophthora fragariae var. rubi Phytophthora fragariae var. rubi Phytophthora rubi is the principal cause of root rot in red raspberry. Originally classified as a variant of P. fragariae some now consider it a distinct species . This organism has been isolated from raspberries in Europe, North America , Chile and other many countries around the world... Phytophthora megasperma Phytophthora megasperma Phytophthora megasperma is a plant pathogen.- External links :* *... Phytophthora cactorum Phytophthora cactorum Phytophthora cactorum is a plant pathogen that causes root rot on rhododendron and many other species.- External links :* *... Phytophthora citricola Phytophthora citricola Phytophthora citricola is a plant pathogen. It was first described by Sawada in 1927 when it was isolated from orange trees in present-day Taiwan. It has since been found causing disease on a wide variety of plants and is considered to be cosmopolitan.... Phytophthora cryptogea Phytophthora cryptogea Phytophthora cryptogea is a plant pathogen.- External links :**... Phytophthora drechsleri Phytophthora drechsleri Phytophthora drechsleri is a plant pathogen.- External links :**... Phytophthora cambivora Phytophthora cambivora Phytophthora cambivora is a plant pathogen that causes Ink disease in European Chestnut trees . Ink disease, also caused by Phytophthora cinnamomi, is thought to have been present in Europe since the 18th century and causes chestnut trees to wilt and die, major epidemics occurred during the 19th... Phytophthora cinnamomi Phytophthora cinnamomi Phytophthora cinnamomi is a soil-borne water mould that produces an infection which causes a condition in plants called root rot or dieback. The plant pathogen is one of the world's most invasive species and is present in over 70 countries from around the world.- Life cycle and effects on plants :P... Phytophthora erythroseptica Phytophthora erythroseptica Phytophthora erythroseptica is a plant pathogen. It infects potatoes causing their tubers to turn pink and damages leaves. It also infects tulips damaging their leaves and shoots.- External links :... |
Post harvest soft rot = leak disease | Rhizopus stolonifer Rhizopus nigricans Rhizopus nigricans Rhizopus nigricans is a fungus commonly known as bread mold and is the most common species of Rhizopus. It is found on old food and in soils and even in children's sandboxes. The genus contains some 50 species and bread mold is sometimes confused with species of Mucor or other species of Rhizopus... Mucor piriformis Mucor piriformis Mucor piriformis is a plant pathogen that causes a soft rot of several fruits known as Mucor rot. Infection of its host fruits, such as apples and pears takes place post-harvest.- External links :... |
Powdery mildew | Sphaerotheca macularis |
Raspberry leaf spot | Sphaerulina rubi Sphaerulina rubi Sphaerulina rubi is a fungal plant pathogen.- External links :* *... Septoria darrowii Septoria darrowii Septoria darrowii is a species of fungus in the Mycosphaerellaceae family. It is a plant pathogen.... [anamorph] Cylindrosporium rubi Cylindrosporium rubi Cylindrosporium rubi is a plant pathogen.- External links :... |
Root rot | Collybia dryophila Cylindrocarpon destructans Fusarium Fusarium Fusarium is a large genus of filamentous fungi widely distributed in soil and in association with plants. Most species are harmless saprobes, and are relatively abundant members of the soil microbial community. Some species produce mycotoxins in cereal crops that can affect human and animal health... spp. Helicobasidium brebissonii Helicobasidium brebissonii Helicobasidium purpureum is a plant pathogen. It contains 3 variations.-Variations:*Helicobasidium purpureum var. barlae Bres. 1909*Helicobasidium purpureum var. orientalePat. 1920*Helicobasidium purpureum var. purpureum Pat. 1885... Rhizoctonia crocorum [anamorph] Phymatotrichopsis omnivora Phymatotrichum omnivorum Pythium Pythium Pythium is a genus of parasitic oomycete. Most species are plant parasites, but Pythium insidiosum is an important pathogen of animals... spp. Rhizoctonia Rhizoctonia Rhizoctonia is a genus of anamorphic fungi in the order Cantharellales. Species do not produce spores, but are composed of hyphae and sclerotia and are asexual states of fungi in the genus Thanatephorus. Rhizoctonia species are saprotrophic, but are also facultative plant pathogens, causing... spp. |
Rosette = double blossom | Cercosporella rubi Cercosporella rubi Cercosporella rubi is a plant pathogen which causes rossette disease of blackberry. The disease is a particular problem in the southeast of the USA. The disease causes witch's broom symptoms as well as double blossoms, and can severely reduce yield.... |
Septoria leaf spot (blackberry) = cane and leaf spot | Septoria rubi Mycosphaerella rubi Mycosphaerella rubi Mycosphaerella rubi is a fungal plant pathogen.-External links:* *... [teleomorph] |
Silver leaf | Chondrostereum purpureum Chondrostereum purpureum Silver leaf is a fungal disease of trees caused by the fungus plant pathogen Chondrostereum purpureum. It attacks most species of the rose family Rosaceae, particularly the genus Prunus. The disease is progressive and often fatal. The common name is taken from the progressive silvering of leaves on... Stereum purpureum |
Spur blight | Didymella applanata Didymella applanata Didymella applanata is a plant pathogen.- External links :**... Phoma Phoma Phoma is a genus of common coelomycetous soil fungi. It contains many plant pathogenic species. About 140 Phoma taxa have been defined and recognized which may be divided into two large groups: plurivorous fungi, generally saprobic or weakly parasitic, mainly from temperate regions in Eurasia, but... sp. [anamorph] |
Stamen blight | Hapalosphaeria deformans Hapalosphaeria deformans Hapalosphaeria deformans is an ascomycete fungus that is a plant pathogen.- External links :**... |
Sydowiella cane canker | Sydowiella depressula Sydowiella depressula Sydowiella depressula is a plant pathogen.- External links :* *... Phragmidium imitans |
Verticillium wilt | Verticillium albo-atrum Verticillium albo-atrum Verticillium albo-atrum is a plant pathogen.-External links:* *... Verticillium dahliae Verticillium dahliae Verticillium dahliae is a fungal plant pathogen. It causes verticillium wilt in many plant species, causing leaves to curl and discolor. It may cause death in some plants.- External links :* *... |
White root rot | Vararia Vararia Vararia is a genus of fungi in the Lachnocladiaceae family. The genus contains 54 species that collectively have a widespread distribution.-External links:* at Index Fungorum... sp. |
Nematodes, parasitic
Nematodes, parasitic |
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Dagger, American nematode | Xiphinema americanum Xiphinema americanum Xiphinema americanum is a plant pathogenic nematode. It is one of many species that belongs to the genus Xiphinema. It was first described by N. A. Cobb in 1913, who found it on both sides of the United States on the roots of grass, corn, and citrus trees. Not only is Xiphinema americanum known to... |
Dagger nematode | Xiphinema bakeri Xiphinema bakeri Xiphinema bakeri is a plant pathogenic nematode.- External links :*... Xiphinema rivesi Xiphinema rivesi Xiphinema rivesi is a plant pathogenic nematode.- External links :*... |
Root-lesion nematode | Pratylenchus Pratylenchus Pratylenchus is a genus of plant-parasitic nematodes responsible for root lesions on many plant hosts in temperate regions around the world. Lesion nematodes are migratory endoparasites that enter the host root for feeding and reproduction and move freely through or out of the root tissue. They do... spp. Pratylenchus penetrans Pratylenchus penetrans Pratylenchus penetrans is a plant pathogenic nematode.- External links :*... |
Needle nematode | Longidorus spp. |
Virus and viruslike agents
Virus and viruslike agents |
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Mosaic or ringspot | genus Ilarvirus Ilarvirus Ilarvirus is a genus of viruses.-External links:*... , Apple mosaic virus Apple mosaic virus Apple mosaic virus is a plant pathogenic virus of the family Bromoviridae.-External links:**... (ApMV) |
Tobacco ringspot in Rubus | genus Nepovirus Nepovirus Nepoviruses or nematode transmitted polyhedral viruses, are a genus of plant viruses of the family Secoviridae and subfamily Comovirinae... , Tobacco ringspot virus Tobacco ringspot virus Tobacco ringspot virus is a plant pathogenic virus in the plant virus family Comoviridae. It is the type species of the Comoviridae Genus Nepovirus. Nepoviruses are be transmitted between plants by nematodes... (TRSV) |
Black raspberry streak | A graft-transmissible agent(s) of unknown identity |
Blackberry calico | genus Carlavirus Carlavirus Carlavirus, also known as "Carnation latent virus group" is a genus of virus that infects plants. It contains thirty-five described species and twenty-nine tentative species..-Virology:... , Blackberry calico virus (BCV) in North America. genus Potexvirus Potexvirus Potexvirus is a genus of plant pathogenic viruses, belonging to the family Alphaflexiviridae and the order Tymovirales. Transmission occurs mechanically, with viruses in this genus causing mosaic and ringspot symptoms.... , Wineberry latent virus Wineberry latent virus Wineberry latent virus is a plant pathogenic virus of the family Flexiviridae.-External links:**... (WLV) upon inoculation in the U.K. |
Bramble yellow mosaic | genus Potyvirus Potyvirus Potyviruses infect plants and belong to the family Potyviridae. The genus is named after the type virus - Potato Virus Y.The Potyvirus, like the Begomoviruses, have ~30% of the currently known plant viruses and have at least 180 definitive and possible members... , Bramble yellow mosaic virus Bramble yellow mosaic virus Bramble yellow mosaic virus is a plant pathogenic virus of the family Potyviridae.-External links:**... (BrmYMV) |
European decline | genus Nepovirus Nepovirus Nepoviruses or nematode transmitted polyhedral viruses, are a genus of plant viruses of the family Secoviridae and subfamily Comovirinae... , Cherry leaf roll virus Cherry leaf roll virus Cherry leaf roll virus is a plant pathogenic virus of the subfamily Comovirinae, family Secoviridae, order Picornavirales.-External links:***ICTV Virus Taxonomy 2009 *UniProt Taxonomy... (CLRV) |
Green blotch | genus Cucumovirus Cucumovirus Cucumovirus is a genus of virus that infects plants. It is a member of the Bromoviridae family. It contains three described species, the cucumber mosaic virus, peanut stunt virus and tomato aspermy virus.... , Cucumber mosaic virus Cucumber mosaic virus Cucumber mosaic virus is a plant pathogenic virus in the family Bromoviridae.It is the type member of the plant virus genus, Cucumovirus. This virus has a worldwide distribution and a very wide host range. In fact it has the reputation of having the widest host range of any known plant virus... (CMV) |
Latent virus, North America | genus Ilarvirus Ilarvirus Ilarvirus is a genus of viruses.-External links:*... , Tobacco streak virus Tobacco streak virus Tobacco streak virus is a plant pathogenic virus of the family Bromoviridae.-External links:**- Affected organisms :* Asparagus, Asparagus officinalis* Faba bean, Vicia faba* Cotton, Gossypium herbaceum* Sun flower* Groundnut... (Rubus strain) (TSV-R) |
Latent virus, Europe | genus Potexvirus Potexvirus Potexvirus is a genus of plant pathogenic viruses, belonging to the family Alphaflexiviridae and the order Tymovirales. Transmission occurs mechanically, with viruses in this genus causing mosaic and ringspot symptoms.... , Wineberry latent virus Wineberry latent virus Wineberry latent virus is a plant pathogenic virus of the family Flexiviridae.-External links:**... (WLV) |
North American raspberry decline | genus Nepovirus Nepovirus Nepoviruses or nematode transmitted polyhedral viruses, are a genus of plant viruses of the family Secoviridae and subfamily Comovirinae... , Tomato ringspot virus Tomato ringspot virus Tomato ringspot virus is a plant pathogenic virus of the family Comoviridae. It affects tomatoes.-External links:* Descriptions of Plant Viruses: **... (TomRSV) or genus Nepovirus Nepovirus Nepoviruses or nematode transmitted polyhedral viruses, are a genus of plant viruses of the family Secoviridae and subfamily Comovirinae... , Cherry rasp leaf virus Cherry rasp leaf virus Cherry rasp leaf virus is a plant pathogenic virus of the order Picornavirales, family Secoviridae, subfamily Comovirinae, genus Cheravirus.- Symptoms :... (CRLV) |
Raspberry bushy dwarf | genus Ideaovirus, Raspberry bushy dwarf virus Raspberry bushy dwarf virus Raspberry bushy dwarf virus is a plant pathogenic virus.- Causes :RBDV is usually transmitted through infected pollen. Insects that feed from infected bushes are also capable of spreading the infected pollen.- Symptoms :... (RBDV) |
Raspberry mosaic | In North America caused by complexes of: Rubus yellow net virus Rubus yellow net virus Rubus yellow net virus is a plant pathogenic virus of the family Caulimoviridae.-External links:**... (RYNV); black raspberry necrosis virus Black raspberry necrosis virus Black raspberry necrosis virus is a plant pathogenic virus.-External links:**... (BRNV); and an unnamed isometric virus similar to raspberry leaf mottle virus (RLMV) |
Raspberry ringspot | Caused by either genus Nepovirus Nepovirus Nepoviruses or nematode transmitted polyhedral viruses, are a genus of plant viruses of the family Secoviridae and subfamily Comovirinae... , Raspberry ringspot virus Raspberry ringspot virus Raspberry ringspot virus is a plant pathogenic virus of the family Comoviridae.-External links:**... (RRSV) or genus Nepovirus Nepovirus Nepoviruses or nematode transmitted polyhedral viruses, are a genus of plant viruses of the family Secoviridae and subfamily Comovirinae... tomato black ring virus Tomato black ring virus Tomato black ring virus is a plant pathogenic virus of the family Comoviridae.-External links:**... (TBRV) |
Raspberry vein chlorosis | Raspberry vein chlorosis virus Raspberry vein chlorosis virus Raspberry vein chlorosis virus is a plant pathogenic virus of the family Rhabdoviridae.-External links:**... (RVCV) |
Raspberry veinbanding mosaic | Caused by RYNV + RLMV; additionally raspberry leaf spot virus (RLSV), uncharacterized, may occur with this complex |
Raspberry yellow dwarf | Caused by either genus Nepovirus Nepovirus Nepoviruses or nematode transmitted polyhedral viruses, are a genus of plant viruses of the family Secoviridae and subfamily Comovirinae... , Arabis mosaic virus Arabis mosaic virus Arabis mosaic virus is a plant pathogenic virus of the family Comoviridae.-External links:**... (ArMV) or genus Nepovirus Nepovirus Nepoviruses or nematode transmitted polyhedral viruses, are a genus of plant viruses of the family Secoviridae and subfamily Comovirinae... , Strawberry latent ringspot virus Strawberry latent ringspot virus Strawberry latent ringspot virus is a plant pathogenic virus in an Unassigned family. Its entry in the ICTV database indicates this virus is a Sadwavirus but current literature indicates that it may be a Cheravirus.-References:... (SLRV) |
Raspberry yellow spot | Raspberry yellow spot (virus-like agent of unknown relationship) |
Thimbleberry ringspot | genus Ilarvirus Ilarvirus Ilarvirus is a genus of viruses.-External links:*... , Apple mosaic virus Apple mosaic virus Apple mosaic virus is a plant pathogenic virus of the family Bromoviridae.-External links:**... |
Phytoplasmla and spiroplasmal diseases
Phytoplasmla and spiroplasmal diseases |
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Black raspberry witches’-broom | Phytoplasma |
Rubus stunt | Rubus stunt phytoplasma |
Miscellaneous diseases and disorders
Miscellaneous diseases and disorders |
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Alpine mosaic in Rubus | Alpine mosaic agent, (a graft-transmissible agent of unknown identity) |
Blackberry sterility | Cause unknown |
Brown berry disease (of black raspberry) | Cause unknown |
Crumbly berries | Various causes: poor pollination, genetic, virus, insect, nutrition, winter injury, water relations |
Dryberry disease (loganberry) | Phyllocoptes gracilis (dryberry mite) and Rhizoctonia rubi |
Fasciation | Cause unknown |
Midge blight | Phoma macrostoma var. macrostoma Fusarium culmorum Alternaria spp. colonization following feeding by Resseliella theoboldi (raspberry cane midge) |
Raspberry leaf curl | Raspberry leaf curl uncharacterized agent(s) |
Redberry disease | Acalitus essigi (redberry mite) |
Seedborne dsRNA in wild raspberry | dsRNA of mol. wt. typical of plant viruses, but not associated with host symptoms or virus-like particles |