Medium-billed Prion
Encyclopedia
Salvin's Prion, Pachyptila salvini, also known as Medium-billed Prion, is a species of seabird
Seabird
Seabirds are birds that have adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same environmental problems and feeding niches have resulted in similar adaptations...

 in the petrel
Petrel
Petrels are tube-nosed seabirds in the bird order Procellariiformes. The common name does not indicate relationship beyond that point, as "petrels" occur in three of the four families within that group...

 family Procellariidae
Procellariidae
The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the prions, and the shearwaters. This family is part of the bird order Procellariiformes , which also includes the albatrosses, the storm-petrels, and the diving petrels.The procellariids are...

.

Taxonomy

Salvin's Prion is a member of the Pachyptila
Pachyptila
Pachyptila is a genus from the family Procellariidae and the Procellariiformes order. The members of this genus and the Blue Petrel form a sub-group called Prions.-Etymology:...

genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

, and along with the Blue Petrel
Blue Petrel
The Blue Petrel is a small seabird in the family Procellariidae. This small petrel is the only member of the genus Halobaena but is closely allied to the prions.-Taxonomy:...

, they make up the prions
Prion (bird)
The Prions are small petrels in the genera Pachyptila and Halobaena. They form one of the four groups within the Procellariidae , along with the gadfly petrels, shearwaters and fulmarine petrels....

. They in turn are members of the Procellariidae
Procellariidae
The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the prions, and the shearwaters. This family is part of the bird order Procellariiformes , which also includes the albatrosses, the storm-petrels, and the diving petrels.The procellariids are...

 family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

, and the Procellariiformes
Procellariiformes
Procellariiformes is an order of seabirds that comprises four families: the albatrosses, petrels and shearwaters, storm petrels, and diving petrels...

 order
Order (biology)
In scientific classification used in biology, the order is# a taxonomic rank used in the classification of organisms. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, and species, with order fitting in between class and family...

. The prions are small and typically eat just zooplankton
Zooplankton
Zooplankton are heterotrophic plankton. Plankton are organisms drifting in oceans, seas, and bodies of fresh water. The word "zooplankton" is derived from the Greek zoon , meaning "animal", and , meaning "wanderer" or "drifter"...

; however as a member of the Procellariiformes, they share certain identifying features. First, they have nasal passages that attach to the upper bill called naricorns. Although the nostrils on the Prion are on top of the upper bill. The bills of Procellariiformes are also unique in that they are split into between 7 and 9 horny plates. They produce a stomach oil made up of wax ester
Wax ester
An ester of a fatty acid and a long-chain alcohol. These oils have similar properties as triglycerides, but are indigestible. They are found in some fish such as orange roughy, oilfish, escolar, black oreo, smooth oreo and other deep water fish. They are also present in marine copepods. Wax...

s and triglycerides that is stored in the proventriculus
Proventriculus
The proventriculus is part of the digestive system of birds, invertebrates and insects.-Birds:The proventriculus is a standard part of avian anatomy...

. This is used against predators as well as an energy rich food source for chicks and for the adults during their long flights. Finally, they also have a salt gland that is situated above the nasal passage and helps desalinate their bodies, due to the high amount of ocean water that they imbibe. It excretes a high saline solution from their nose.

Subspecies

Salvin's Prion has two subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

:
  • Pachyptila salvini salvini, the nominate subspecies, breeds on Prince Edward Island
    Prince Edward Islands
    The Prince Edward Islands are two small islands in the sub-antarctic Indian Ocean that are part of South Africa. The islands, named Marion Island and Prince Edward Island, are located at ....

     and the Crozet Islands
    Crozet Islands
    The Crozet Islands are a sub-antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarctic Lands.-Geography:...

  • Pachyptila salvini macgillivray breeds on St. Paul Island
    Île Saint-Paul
    Île Saint-Paul is an island forming part of the French Southern and Antarctic Lands in the Indian Ocean, with an area of . It is located about southwest of the larger Île Amsterdam, and south of Réunion...

     and Amsterdam Island

Etymology

The name Pachyptila comes from the Greek
Greek language
Greek is an independent branch of the Indo-European family of languages. Native to the southern Balkans, it has the longest documented history of any Indo-European language, spanning 34 centuries of written records. Its writing system has been the Greek alphabet for the majority of its history;...

 words pakhus and ptilon. Pakhus means "thick" or "stout" and ptilon means "a feather". Also from the Greek language, Prion comes from the word priōn meaning "a saw", which is in reference to its serrated edges of its bill
Beak
The beak, bill or rostrum is an external anatomical structure of birds which is used for eating and for grooming, manipulating objects, killing prey, fighting, probing for food, courtship and feeding young...

. The species is named for the British
United Kingdom
The United Kingdom of Great Britain and Northern IrelandIn the United Kingdom and Dependencies, other languages have been officially recognised as legitimate autochthonous languages under the European Charter for Regional or Minority Languages...

 ornithologist
Ornithology
Ornithology is a branch of zoology that concerns the study of birds. Several aspects of ornithology differ from related disciplines, due partly to the high visibility and the aesthetic appeal of birds...

 Osbert Salvin
Osbert Salvin
Osbert Salvin FRS was an English naturalist, best known for co-authoring Biologia Centrali-Americana with Frederick DuCane Godman. This was a 52 volume encyclopedia on the natural history of Central America....

.

Description

Salvin's Prion is a small 29 cm (11.4 in) petrel with grey and white plumage
Plumage
Plumage refers both to the layer of feathers that cover a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage vary between species and subspecies and can also vary between different age classes, sexes, and season. Within species there can also be a...

, and a blue bill. Like the Broad-billed Prion
Broad-billed Prion
The Broad-billed Prion, Pachyptila vittata, is a small seabird, but the largest Prion, with grey upperparts plumage, and white underparts. It has many other names that have been used such as Blue-billed Dove-petrel, Broad-billed Dove-petrel, Long-billed Prion, Common Prion, Icebird, and...

 it has lamellae in its bill
Beak
The beak, bill or rostrum is an external anatomical structure of birds which is used for eating and for grooming, manipulating objects, killing prey, fighting, probing for food, courtship and feeding young...

 in order to filter seawater for food.

Breeding

This small prion
Prion (bird)
The Prions are small petrels in the genera Pachyptila and Halobaena. They form one of the four groups within the Procellariidae , along with the gadfly petrels, shearwaters and fulmarine petrels....

 breeds colonially on a number of subantarctic
Subantarctic
The Subantarctic is a region in the southern hemisphere, located immediately north of the Antarctic region. This translates roughly to a latitude of between 46° – 60° south of the Equator. The subantarctic region includes many islands in the southern parts of the Indian Ocean, Atlantic Ocean and...

 islands in the southern Indian Ocean
Indian Ocean
The Indian Ocean is the third largest of the world's oceanic divisions, covering approximately 20% of the water on the Earth's surface. It is bounded on the north by the Indian Subcontinent and Arabian Peninsula ; on the west by eastern Africa; on the east by Indochina, the Sunda Islands, and...

. The colonies of Medium-billed Prions are attended nocturnally in order to avoid predation by skua
Skua
The skuas are a group of seabirds with about seven species forming the family Stercorariidae and the genus Stercorarius. The three smaller skuas are called jaegers in North America....

s. The nests are concealed in burrows usually dug into soil. Nests are attended regularly for several months prior to breeding. A single egg is laid in November or early December, which is incubated
Avian incubation
Incubation refers to the process by which certain oviparous animals hatch their eggs, and to the development of the embryo within the egg. The most vital factor of incubation is the constant temperature required for its development over a specific period. Especially in domestic fowl, the act of...

 for around 50 days. Both parents share the incubation duties and feed the chick once it is hatched. The chicks fledge
Fledge
Fledge is the stage in a young bird's life when the feathers and wing muscles are sufficiently developed for flight. It also describes the act of a chick's parents raising it to a fully grown state...

 around 60 days after hatching.

Feeding

The main components of its diet are amphipods and krill
Krill
Krill is the common name given to the order Euphausiacea of shrimp-like marine crustaceans. Also known as euphausiids, these small invertebrates are found in all oceans of the world...

, although it will also take fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...

 and squid
Squid
Squid are cephalopods of the order Teuthida, which comprises around 300 species. Like all other cephalopods, squid have a distinct head, bilateral symmetry, a mantle, and arms. Squid, like cuttlefish, have eight arms arranged in pairs and two, usually longer, tentacles...

. In addition to filter feeding food is obtained by seizing and hydroplaning.

Range and habitat

Salvin's Prion breeds principally on Île aux Cochons in the Crozet Islands
Crozet Islands
The Crozet Islands are a sub-antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarctic Lands.-Geography:...

, where four million pairs are thought to breed. Other breeding colonies include Prince Edward Island
Prince Edward Island
Prince Edward Island is a Canadian province consisting of an island of the same name, as well as other islands. The maritime province is the smallest in the nation in both land area and population...

, St Paul Island
St Paul Island
Saint Paul Island can refer to:* Île Saint-Paul, a small island in the French Southern Territories* Saint Paul Island , United States* St. Paul Island , Canada* Saint Peter and Saint Paul Archipelago, Brazil* St Paul's Island, Malta...

 and Amsterdam Island. At sea they range from South Africa
South Africa
The Republic of South Africa is a country in southern Africa. Located at the southern tip of Africa, it is divided into nine provinces, with of coastline on the Atlantic and Indian oceans...

 eastwards to New Zealand
New Zealand
New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

.

Conservation

Salvin's Prion is not considered threatened. Although numbers have declined on some islands where rat
Rat
Rats are various medium-sized, long-tailed rodents of the superfamily Muroidea. "True rats" are members of the genus Rattus, the most important of which to humans are the black rat, Rattus rattus, and the brown rat, Rattus norvegicus...

s and feral cat
Feral cat
A feral cat is a descendant of a domesticated cat that has returned to the wild. It is distinguished from a stray cat, which is a pet cat that has been lost or abandoned, while feral cats are born in the wild; the offspring of a stray cat can be considered feral if born in the wild.In many parts of...

s have been introduced
Introduced species
An introduced species — or neozoon, alien, exotic, non-indigenous, or non-native species, or simply an introduction, is a species living outside its indigenous or native distributional range, and has arrived in an ecosystem or plant community by human activity, either deliberate or accidental...

, the world population is estimated at around 12 million birds. Consequently, they are given a classification of Least Concern
Least Concern
Least Concern is an IUCN category assigned to extant taxon or lower taxa which have been evaluated but do not qualify for any other category. As such they do not qualify as threatened, Near Threatened, or Conservation Dependent...

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