Common names: pine snake, pinesnake, common pine snake, more.

Pituophis melanoleucus is a harmless colubrid

Colubrid

A colubrid is a member of the snake family Colubridae. This broad classification of snakes includes about two-thirds of all snake species on earth. The earliest species of the snake family date back to the Oligocene epoch. With 304 genera and 1,938 species, Colubridae is the largest snake family...

 species

Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 found in the United States

United States

The United States of America is a federal constitutional republic comprising fifty states and a federal district...

. Three subspecies

Subspecies

Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

 are currently recognized, including the nominate subspecies described here.

Description

Adults are large, growing to 48-100 inches (122-254 cm) in length, and are powerfully built. The head is small and somewhat pointed with an enlarged rostral scale that extends upward between the internasal scales

Internasal scales

In snakes, the internasal scales are those on top of the head between the scales that surround the nostrils. They are usually paired and situated just behind the rostral....

. There are usually 4 prefrontal scales

Prefrontal scales

The prefrontal scales on snakes and other reptiles are the scales connected to the frontals towards the tip of the snout which are in contact with the internasals....

. At midbody there are 27-37 rows of keeled

Keeled scales

Keeled scales refer to reptile scales that, rather than being smooth, have a ridge down the center that may or may not extend to the tip of the scale, making them rough to the touch...

 dorsal scales

Dorsal scales

In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales.When counting dorsal scales, numbers are often given for three points along the body, for example 19:21:17...

. The anal plate is single. The color pattern consists of a light ground color overlaid with black, brown or reddish-brown blotches.

Common names

Pine snake, pinesnake, common pine snake, bull snake, black and white snake, carpet snake, chicken snake, kawaiiest snakeicus , common bull snake, eastern bull snake, eastern pine snake, horn(ed) snake, New Jersey pine snake, North American pine snake, northern pine snake, pilot snake, white gopher snake.

Geographic range

Found in the United States

United States

The United States of America is a federal constitutional republic comprising fifty states and a federal district...

 in Washington, Oregon

Oregon

Oregon is a state in the Pacific Northwest region of the United States. It is located on the Pacific coast, with Washington to the north, California to the south, Nevada on the southeast and Idaho to the east. The Columbia and Snake rivers delineate much of Oregon's northern and eastern...

, Montana

Montana

Montana is a state in the Western United States. The western third of Montana contains numerous mountain ranges. Smaller, "island ranges" are found in the central third of the state, for a total of 77 named ranges of the Rocky Mountains. This geographical fact is reflected in the state's name,...

, Wyoming

Wyoming

Wyoming is a state in the mountain region of the Western United States. The western two thirds of the state is covered mostly with the mountain ranges and rangelands in the foothills of the Eastern Rocky Mountains, while the eastern third of the state is high elevation prairie known as the High...

, California

California

California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

, Nevada

Nevada

Nevada is a state in the western, mountain west, and southwestern regions of the United States. With an area of and a population of about 2.7 million, it is the 7th-largest and 35th-most populous state. Over two-thirds of Nevada's people live in the Las Vegas metropolitan area, which contains its...

, Arizona

Arizona

Arizona ; is a state located in the southwestern region of the United States. It is also part of the western United States and the mountain west. The capital and largest city is Phoenix...

, Utah

Utah

Utah is a state in the Western United States. It was the 45th state to join the Union, on January 4, 1896. Approximately 80% of Utah's 2,763,885 people live along the Wasatch Front, centering on Salt Lake City. This leaves vast expanses of the state nearly uninhabited, making the population the...

, southwestern North Dakota

North Dakota

North Dakota is a state located in the Midwestern region of the United States of America, along the Canadian border. The state is bordered by Canada to the north, Minnesota to the east, South Dakota to the south and Montana to the west. North Dakota is the 19th-largest state by area in the U.S....

, South Dakota

South Dakota

South Dakota is a state located in the Midwestern region of the United States. It is named after the Lakota and Dakota Sioux American Indian tribes. Once a part of Dakota Territory, South Dakota became a state on November 2, 1889. The state has an area of and an estimated population of just over...

, Nebraska

Nebraska

Nebraska is a state on the Great Plains of the Midwestern United States. The state's capital is Lincoln and its largest city is Omaha, on the Missouri River....

, Oklahoma

Oklahoma

Oklahoma is a state located in the South Central region of the United States of America. With an estimated 3,751,351 residents as of the 2010 census and a land area of 68,667 square miles , Oklahoma is the 28th most populous and 20th-largest state...

, New Mexico

New Mexico

New Mexico is a state located in the southwest and western regions of the United States. New Mexico is also usually considered one of the Mountain States. With a population density of 16 per square mile, New Mexico is the sixth-most sparsely inhabited U.S...

, Texas

Texas

Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

, Minnesota

Minnesota

Minnesota is a U.S. state located in the Midwestern United States. The twelfth largest state of the U.S., it is the twenty-first most populous, with 5.3 million residents. Minnesota was carved out of the eastern half of the Minnesota Territory and admitted to the Union as the thirty-second state...

, Iowa

Iowa

Iowa is a state located in the Midwestern United States, an area often referred to as the "American Heartland". It derives its name from the Ioway people, one of the many American Indian tribes that occupied the state at the time of European exploration. Iowa was a part of the French colony of New...

, Louisiana

Louisiana

Louisiana is a state located in the southern region of the United States of America. Its capital is Baton Rouge and largest city is New Orleans. Louisiana is the only state in the U.S. with political subdivisions termed parishes, which are local governments equivalent to counties...

, Illinois

Illinois

Illinois is the fifth-most populous state of the United States of America, and is often noted for being a microcosm of the entire country. With Chicago in the northeast, small industrial cities and great agricultural productivity in central and northern Illinois, and natural resources like coal,...

, Wisconsin

Wisconsin

Wisconsin is a U.S. state located in the north-central United States and is part of the Midwest. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake Michigan to the east, Michigan to the northeast, and Lake Superior to the north. Wisconsin's capital is...

, Indiana

Indiana

Indiana is a US state, admitted to the United States as the 19th on December 11, 1816. It is located in the Midwestern United States and Great Lakes Region. With 6,483,802 residents, the state is ranked 15th in population and 16th in population density. Indiana is ranked 38th in land area and is...

, Kentucky

Kentucky

The Commonwealth of Kentucky is a state located in the East Central United States of America. As classified by the United States Census Bureau, Kentucky is a Southern state, more specifically in the East South Central region. Kentucky is one of four U.S. states constituted as a commonwealth...

, Tennessee

Tennessee

Tennessee is a U.S. state located in the Southeastern United States. It has a population of 6,346,105, making it the nation's 17th-largest state by population, and covers , making it the 36th-largest by total land area...

, Virginia

Virginia

The Commonwealth of Virginia , is a U.S. state on the Atlantic Coast of the Southern United States. Virginia is nicknamed the "Old Dominion" and sometimes the "Mother of Presidents" after the eight U.S. presidents born there...

, North Carolina

North Carolina

North Carolina is a state located in the southeastern United States. The state borders South Carolina and Georgia to the south, Tennessee to the west and Virginia to the north. North Carolina contains 100 counties. Its capital is Raleigh, and its largest city is Charlotte...

, South Carolina

South Carolina

South Carolina is a state in the Deep South of the United States that borders Georgia to the south, North Carolina to the north, and the Atlantic Ocean to the east. Originally part of the Province of Carolina, the Province of South Carolina was one of the 13 colonies that declared independence...

, Georgia

Georgia (U.S. state)

Georgia is a state located in the southeastern United States. It was established in 1732, the last of the original Thirteen Colonies. The state is named after King George II of Great Britain. Georgia was the fourth state to ratify the United States Constitution, on January 2, 1788...

, Alabama

Alabama

Alabama is a state located in the southeastern region of the United States. It is bordered by Tennessee to the north, Georgia to the east, Florida and the Gulf of Mexico to the south, and Mississippi to the west. Alabama ranks 30th in total land area and ranks second in the size of its inland...

, Mississippi

Mississippi

Mississippi is a U.S. state located in the Southern United States. Jackson is the state capital and largest city. The name of the state derives from the Mississippi River, which flows along its western boundary, whose name comes from the Ojibwe word misi-ziibi...

, Florida

Florida

Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...

 and New Jersey

New Jersey

New Jersey is a state in the Northeastern and Middle Atlantic regions of the United States. , its population was 8,791,894. It is bordered on the north and east by the state of New York, on the southeast and south by the Atlantic Ocean, on the west by Pennsylvania and on the southwest by Delaware...

. It also occurs in southwestern Canada

Canada

Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...

 and in Mexico

Mexico

The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

.

The range for P. m. melanoleucus extends from southern North Carolina west through South Carolina to northern Georgia, eastern Tennessee, southeastern Kentucky and south into Alabama. Disjunct populations exist in southern New Jersey, west-central Virginia, adjacent West Virginia, central Kentucky and southwestern Tennessee. This subspecies intergrades with P. m. mugitus in South Carolina, Georgia, and Alabama.

Conservation status

This species is classified as Least Concern (LC) on the IUCN

World Conservation Union

The International Union for Conservation of Nature and Natural Resources is an international organization dedicated to finding "pragmatic solutions to our most pressing environment and development challenges." The organization publishes the IUCN Red List, compiling information from a network of...

 Red List of Threatened Species

IUCN Red List

The IUCN Red List of Threatened Species , founded in 1963, is the world's most comprehensive inventory of the global conservation status of biological species. The International Union for Conservation of Nature is the world's main authority on the conservation status of species...

 (v3.1, 2001). Species are listed as such due to their wide distribution, presumed large population, or because it is unlikely to be declining fast enough to qualify for listing in a more threatened category. The population trend is down. Year assessed: 2007.

Habitat

Prefers pine flatwoods, sandy pine-oak woodlands, prairies, cultivated fields, open brushland, rocky desert and chaparral. Occurs from sea level to an elevation of 9,000 feet (2,750 m).

Behavior

When disturbed, it will often hiss loudly, sometimes flattening its head, vibrate its tail and eventually strike at an intruder. Some species will use 'Cloacal Popping' as a defense mechanism. To make the sound, the snakes contract their cloacal sphincter, forcing air (and any other material that happens to be there) out. It is an extremely foul smell that sometimes scares away predators.

Feeding

Prey includes rats, mice, moles, other small mammals and bird's eggs. According to Mehrtens (1987), it often enters rodent burrows in search of a meal. In these cases, multiple kills are frequent with the snake pressing the mice against the walls of the burrow.

Reproduction

After mating has taken place in the spring, clutches of 3-24 eggs are laid in June-August. The eggs are deposited in sandy burrows or under large rocks or logs and hatch after 64-79 days of incubation. The eggs are adherent and quite large, up to 66 mm (2⅝ in.) long by 45 mm (1¾ in.) wide. The hatchlings measure 33-45 cm (13-17¾ in.) in total length.

Subspecies

Subspecies	Authority	Common name	Geographic range
P. m. lodingi	Blanchard Frank N. Blanchard Frank Nelson Blanchard was an American herpetologist, and professor of zoology at the University of Michigan. He is credited with identifying several new species, including the Broad-banded Water Snake, Nerodia fasciata confluens, and the Florida King Snake, Lampropeltis getula floridana... , 1924	Black pine snake	From southwestern Alabama to eastern Louisiana.
P. m. melanoleucus	(Daudin François Marie Daudin François Marie Daudin was a French zoologist.With legs paralyzed by childhood disease, he studied of physics and natural history, but ended up being devoted to the latter.Daudin wrote in 1799–1800... , 1803)	Northern pine snake	Southern New Jersey, southern North Carolina west through South Carolina to northern Georgia, eastern Tennessee, southeastern Kentucky and south into Alabama.
P. m. mugitus	Barbour Thomas Barbour Thomas Barbour was an American herpetologist. From 1927 until 1946, he was director of the Museum of Comparative Zoology founded in 1859 by Louis Agassiz at Harvard University in Cambridge, Massachusetts.... , 1921	Florida pine snake	Southern South Carolina to Georgia and southern Florida.

External links

• The Florida Pinesnake: "Pituophis melanoleucus mugitus" - University of Florida Fact Sheet 2009
• "Black Snakes": Identification and Ecology - University of Florida fact sheet
• Northern Pine Snake at The Pituophis Page. Accessed 29 June 2008.
• New Jersey Pine Snake at The State of New Jersey. Accessed 29 June 2008.