Podocarpus
Encyclopedia
Podocarpus is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of conifers, the most numerous and widely distributed of the podocarp family Podocarpaceae
Podocarpaceae
Podocarpaceae is a large family of mainly Southern Hemisphere conifers, comprising about 156 species of evergreen trees and shrubs. It contains 19 genera if Phyllocladus is included and if Manoao and Sundacarpus are recognized....

. The 105 species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of Podocarpus are evergreen
Evergreen
In botany, an evergreen plant is a plant that has leaves in all seasons. This contrasts with deciduous plants, which completely lose their foliage during the winter or dry season.There are many different kinds of evergreen plants, both trees and shrubs...

 shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...

s or tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

s from 1-25 m (rarely to 40 m) in height. The leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 are 0.5-15 cm long, lanceolate to oblong, falcate (sickle-shaped) in some species, with a distinct midrib, and are arranged spirally, though in some species twisted to appear in two horizontal ranks. The cones
Conifer cone
A cone is an organ on plants in the division Pinophyta that contains the reproductive structures. The familiar woody cone is the female cone, which produces seeds. The male cones, which produce pollen, are usually herbaceous and much less conspicuous even at full maturity...

 have two to five fused scales, of which only one, rarely two, are fertile, each fertile scale with one apical seed
Seed
A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...

. At maturity, the scales become berry
Berry
The botanical definition of a berry is a fleshy fruit produced from a single ovary. Grapes are an example. The berry is the most common type of fleshy fruit in which the entire ovary wall ripens into an edible pericarp. They may have one or more carpels with a thin covering and fleshy interiors....

-like, swollen, brightly coloured red to purple and fleshy, and are eaten by bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

s which then disperse the seeds in their droppings. The male (pollen
Pollen
Pollen is a fine to coarse powder containing the microgametophytes of seed plants, which produce the male gametes . Pollen grains have a hard coat that protects the sperm cells during the process of their movement from the stamens to the pistil of flowering plants or from the male cone to the...

) cones are 5-20 mm long, often clustered several together. Many species, though not all, are dioecious
Plant sexuality
Plant sexuality covers the wide variety of sexual reproduction systems found across the plant kingdom. This article describes morphological aspects of sexual reproduction of plants....

.

Podocarpus and the Podocarpaceae were endemic to the ancient supercontinent of Gondwana
Gondwana
In paleogeography, Gondwana , originally Gondwanaland, was the southernmost of two supercontinents that later became parts of the Pangaea supercontinent. It existed from approximately 510 to 180 million years ago . Gondwana is believed to have sutured between ca. 570 and 510 Mya,...

, which broke up into Africa
Africa
Africa is the world's second largest and second most populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area...

, South America
South America
South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

, India
India
India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

, Australia-New Guinea, New Zealand
New Zealand
New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

, and New Caledonia
New Caledonia
New Caledonia is a special collectivity of France located in the southwest Pacific Ocean, east of Australia and about from Metropolitan France. The archipelago, part of the Melanesia subregion, includes the main island of Grande Terre, the Loyalty Islands, the Belep archipelago, the Isle of...

 between 105 and 45 million years ago. Podocarpus is a characteristic tree of the Antarctic flora
Antarctic flora
The Antarctic flora is a distinct community of vascular plants which evolved millions of years ago on the supercontinent of Gondwana, and is now found on several separate areas of the Southern Hemisphere, including southern South America, southernmost Africa, New Zealand, Australia and New Caledonia...

, which originated in the cool, moist climate of southern Gondwana, and elements of the flora survive in the humid temperate regions of the former supercontinent. As the continents drifted north and became drier and hotter, Podocarps and other members of the Antarctic flora generally retreated to humid regions, especially in Australia, where sclerophyll
Sclerophyll
Sclerophyll is the term for a type of vegetation that has hard leaves and short internodes . The word comes from the Greek sclero and phyllon ....

 genera like Acacia
Acacia
Acacia is a genus of shrubs and trees belonging to the subfamily Mimosoideae of the family Fabaceae, first described in Africa by the Swedish botanist Carl Linnaeus in 1773. Many non-Australian species tend to be thorny, whereas the majority of Australian acacias are not...

and Eucalyptus
Eucalyptus
Eucalyptus is a diverse genus of flowering trees in the myrtle family, Myrtaceae. Members of the genus dominate the tree flora of Australia...

became predominant, and the old Antarctic flora retreated to pockets that presently cover only 2% of the continent. As Australia drifted north toward Asia, the collision pushed up the Indonesia
Indonesia
Indonesia , officially the Republic of Indonesia , is a country in Southeast Asia and Oceania. Indonesia is an archipelago comprising approximately 13,000 islands. It has 33 provinces with over 238 million people, and is the world's fourth most populous country. Indonesia is a republic, with an...

n archipelago and the mountains of New Guinea, which allowed podocarp species to hop across the narrow straits into humid Asia, with P. macrophyllus reaching north to southern China
China
Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

 and Japan
Japan
Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

. The flora of Malesia
Malesia
Malesia is a biogeographical region straddling the boundaries of the Indomalaya ecozone and Australasia ecozone, and also a phytogeographical floristic region in the Paleotropical Kingdom.-Floristic province:...

, which includes the Malay peninsula
Malay Peninsula
The Malay Peninsula or Thai-Malay Peninsula is a peninsula in Southeast Asia. The land mass runs approximately north-south and, at its terminus, is the southern-most point of the Asian mainland...

, Indonesia, the Philippines
Philippines
The Philippines , officially known as the Republic of the Philippines , is a country in Southeast Asia in the western Pacific Ocean. To its north across the Luzon Strait lies Taiwan. West across the South China Sea sits Vietnam...

, and New Guinea, is generally derived from Asia but includes many elements of the old Gondwana flora, including several other genera in the Podocarpaceae (Dacrycarpus
Dacrycarpus
Dacrycarpus is a genus of conifers belonging to the Podocarp family, Podocarpaceae. The genus includes nine species of dioecious evergreen trees and shrubs to 55-60 meters in height....

, Dacrydium
Dacrydium
Dacrydium is a genus of conifers belonging to the podocarp family Podocarpaceae. Sixteen species of evergreen dioecious trees and shrubs are presently recognized. The genus was first described by Solander in 1786, and formerly included many more species, which were divided into sections A, B, and C...

, Falcatifolium
Falcatifolium
Falcatifolium is a genus of conifers belonging to the podocarp family Podocarpaceae. The genus includes evergreen dioecious shrubs and large trees of up to 36 meters. Five species are presently recognized...

, Nageia
Nageia
Nageia is a genus of conifers belonging to the podocarp family Podocarpaceae. Nageia includes evergreen shrubs and trees, from one to 54 meters in height. A 2009 treatment of the genus recognized five species. Some authors consider Nageia formosensis to be a separate species from Nageia nagi,...

, Phyllocladus
Phyllocladus
Phyllocladus is a small genus of conifers, now usually treated in the family Podocarpaceae. They are morphologically very distinct from the other genera in that family, and some botanists treat them in a family of their own, the Phyllocladaceae. One molecular phylogenetic analysis found...

,
and the Malesian endemic Sundacarpus
Sundacarpus
Sundacarpus is a genus of conifers containing a single species Sundacarpus amarus, belonging to the family Podocarpaceae. Sundacarpus was designated a genus by C.N.Page in 1989; formerly it had been classified variously as a species of Podocarpus or of Prumnopitys.Sundacarpus is a large evergreen...

), and also Agathis
Agathis
The genus Agathis, commonly known as kauri or dammar, is a relatively small genus of 21 species of evergreen tree. The genus is part of the ancient Araucariaceae family of conifers, a group once widespread during the Jurassic period, but now largely restricted to the Southern Hemisphere except for...

in the Araucariaceae
Araucariaceae
Araucariaceae, commonly referred to as araucarians, is a very ancient family of coniferous trees. It achieved its maximum diversity in the Jurassic and Cretaceous periods, when it was distributed almost worldwide...

.

Classification

There are two subgenera, subgenus Podocarpus and subgenus Foliolatus, distinguished by cone and seed morphology.

Subgenus Podocarpus. Cone not subtended by lanceolate bracts, seed usually with an apical ridge. Distributed in the temperate forests of Tasmania
Tasmania
Tasmania is an Australian island and state. It is south of the continent, separated by Bass Strait. The state includes the island of Tasmania—the 26th largest island in the world—and the surrounding islands. The state has a population of 507,626 , of whom almost half reside in the greater Hobart...

, New Zealand
New Zealand
New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

, southern Chile
Chile
Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...

, with some species extending into the tropical highlands of Africa
Africa
Africa is the world's second largest and second most populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area...

 and the Americas
Americas
The Americas, or America , are lands in the Western hemisphere, also known as the New World. In English, the plural form the Americas is often used to refer to the landmasses of North America and South America with their associated islands and regions, while the singular form America is primarily...

.

Subgenus Foliolatus
. Cone subtended by two lanceolate bracts ("foliola"), seed usually without an apical ridge. Generally tropical and subtropical distribution, concentrated in east and southeast Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

 and Malesia
Malesia
Malesia is a biogeographical region straddling the boundaries of the Indomalaya ecozone and Australasia ecozone, and also a phytogeographical floristic region in the Paleotropical Kingdom.-Floristic province:...

, overlapping with subgenus Podocarpus in northeastern Australia and New Caledonia
New Caledonia
New Caledonia is a special collectivity of France located in the southwest Pacific Ocean, east of Australia and about from Metropolitan France. The archipelago, part of the Melanesia subregion, includes the main island of Grande Terre, the Loyalty Islands, the Belep archipelago, the Isle of...

.

Species in family Podocarpaceae have been reshuffled a number of times based on genetic and physiological evidence, with many species formerly assigned to genus Podocarpus now assigned to other genera. A sequence of classification schemes have moved species between Nageia
Nageia
Nageia is a genus of conifers belonging to the podocarp family Podocarpaceae. Nageia includes evergreen shrubs and trees, from one to 54 meters in height. A 2009 treatment of the genus recognized five species. Some authors consider Nageia formosensis to be a separate species from Nageia nagi,...

and Podocarpus, and in 1969 de Laubenfels divided the huge genus Podocarpus into Dacrycarpus
Dacrycarpus
Dacrycarpus is a genus of conifers belonging to the Podocarp family, Podocarpaceae. The genus includes nine species of dioecious evergreen trees and shrubs to 55-60 meters in height....

, Decussocarpus
(an invalid name he later revised to the valid Nageia
Nageia
Nageia is a genus of conifers belonging to the podocarp family Podocarpaceae. Nageia includes evergreen shrubs and trees, from one to 54 meters in height. A 2009 treatment of the genus recognized five species. Some authors consider Nageia formosensis to be a separate species from Nageia nagi,...

), Prumnopitys
Prumnopitys
Prumnopitys is a genus of conifers belonging to the podocarp family Podocarpaceae. The eight recognised species of Prumnopitys are densely-branched, dioecious evergreen trees up to 40 metres in height. The leaves are similar to those of the yew, strap-shaped, 1-4 cm long and 2-3 mm broad, with a...

, and Podocarpus.

Species
  • Subgenus Podocarpus
    • section Podocarpus (eastern and southern Africa
      Africa
      Africa is the world's second largest and second most populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area...

      )
      • Podocarpus elongatus
        Podocarpus elongatus
        Podocarpus elongatus is a species of conifer in the Podocarpaceae family. Unlike the other Yellowwoods of Southern Africa, Podocarpus elongatus grows as an enormous, dense, multi-stemmed bush....

      • Podocarpus latifolius
        Podocarpus latifolius
        Podocarpus latifolius is a large evergreen tree up to 35 m high and 3 m trunk diameter, in the conifer family Podocarpaceae; it is the type species of the genus Podocarpus....

      • Podocarpus falcatus
    • section Scytopodium (Madagascar
      Madagascar
      The Republic of Madagascar is an island country located in the Indian Ocean off the southeastern coast of Africa...

      , eastern Africa)
      • Podocarpus capuronii
        Podocarpus capuronii
        Podocarpus capuronii is a species of conifer in the Podocarpaceae family.It is found only in Madagascar.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus henkelii
        Podocarpus henkelii
        Podocarpus henkelii is a South African species of conifer in the Podocarpaceae family. It is grown ornamentally in gardens for its strikingly neat, attractive form and its elegant, drooping foliage....

      • Podocarpus humbertii
        Podocarpus humbertii
        Podocarpus humbertii is a species of conifer in the Podocarpaceae family.It is found only in Madagascar.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus madagascariensis
        Podocarpus madagascariensis
        Podocarpus madagascariensis is a species of conifer in the Podocarpaceae family.It is found only in Madagascar.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus rostratus
        Podocarpus rostratus
        Podocarpus rostratus is a species of conifer in the Podocarpaceae family that is found only in Madagascar.-Sources:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

    • section Australis (southeast Australia
      Australia
      Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

      , New Zealand
      New Zealand
      New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

      , New Caledonia
      New Caledonia
      New Caledonia is a special collectivity of France located in the southwest Pacific Ocean, east of Australia and about from Metropolitan France. The archipelago, part of the Melanesia subregion, includes the main island of Grande Terre, the Loyalty Islands, the Belep archipelago, the Isle of...

      , southern Chile
      Chile
      Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...

      )
      • Podocarpus alpinus
        Podocarpus alpinus
        Podocarpus alpinus is a species of conifer in the Podocarpaceae family. It is a shrub about 2-2.5 m tall . It is found only in Australia .-Cultivation and uses:...

      • Podocarpus cunninghamii
        Podocarpus cunninghamii
        Podocarpus cunninghamii is a species of conifer in the Podocarpaceae family, commonly known as Montane totara, thin-bark totara, or Hall's totara. It is also known as Podocarpus hallii can also be found around mountains growing around the base within a forest regionIt is found only in New...

      • Podocarpus gnidioides
        Podocarpus gnidioides
        Podocarpus gnidioides is a species of conifer in the Podocarpaceae family.It is found only in New Caledonia.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus lawrencei
        Podocarpus lawrencei
        Podocarpus lawrencei is a species of podocarp native throughout the Australian high country, from southern Tasmania through to the New South Wales highlands. Common names are Errinundra Plum-pine and Mountain Plum-pine...

      • Podocarpus nivalis
        Podocarpus nivalis
        Podocarpus nivalis is a species of conifer in the Podocarpaceae family.It is found only in New Zealand. It grows in the mountains and subalpine lands from 37° to the far south New Zealand about 46° South Latitude...

      • Podocarpus nubigenus
        Podocarpus nubigenus
        Podocarpus nubigenus is a species of podocarp, endemic to the Valdivian temperate rain forests of southern Chile and adjacent southwestern Argentina. It is the southernmost podocarp in the world, It grows from 38° to Ultima Esperanza , It grows in wet and swampy soils...

      • Podocarpus totara
        Podocarpus totara
        Podocarpus totara is a species of podocarp tree endemic to New Zealand. It grows throughout the North Island and northeastern South Island in lowland, montane and lower subalpine forest at elevations of up to 600 m.-Description:...

    • section Crassiformis (northeast Queensland
      Queensland
      Queensland is a state of Australia, occupying the north-eastern section of the mainland continent. It is bordered by the Northern Territory, South Australia and New South Wales to the west, south-west and south respectively. To the east, Queensland is bordered by the Coral Sea and Pacific Ocean...

      )
      • Podocarpus smithii
        Podocarpus smithii
        Podocarpus smithii is a species of conifer in the Podocarpaceae family.It is found only in Queensland, Australia.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

    • section Capitulatis (central Chile, southern Brazil
      Brazil
      Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

      , the Andes
      Andes
      The Andes is the world's longest continental mountain range. It is a continual range of highlands along the western coast of South America. This range is about long, about to wide , and of an average height of about .Along its length, the Andes is split into several ranges, which are separated...

       from northern Argentina
      Argentina
      Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

       to Ecuador
      Ecuador
      Ecuador , officially the Republic of Ecuador is a representative democratic republic in South America, bordered by Colombia on the north, Peru on the east and south, and by the Pacific Ocean to the west. It is one of only two countries in South America, along with Chile, that do not have a border...

      )
      • Podocarpus glomeratus
        Podocarpus glomeratus
        Podocarpus glomeratus is a species of conifer in the Podocarpaceae family.It is found in Bolivia, Ecuador, and Peru.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus lambertii
        Podocarpus lambertii
        Podocarpus lambertii is a species of conifer in the Podocarpaceae family.It is found in Argentina and Brazil.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus parlatorei
        Podocarpus parlatorei
        Podocarpus parlatorei is a species of Podocarpus, native to Argentina, Bolivia, and Peru.It is a medium-sized to large evergreen tree...

      • Podocarpus salignus
        Podocarpus salignus
        Podocarpus salignus is a species of conifer in the Podocarpaceae family. It is found only in Chile approximately from 35 to 42° South Latitude. Up to 20 m height and 1 m diameter. Straight and cylindrical trunk. The bark is reddish-gray. It is threatened by habitat loss...

      • Podocarpus sellowii
        Podocarpus sellowii
        Podocarpus sellowii is a species of conifer in the Podocarpaceae family.It is found only in Brazil.-Source:* Conifer Specialist Group 2000. .* Downloaded on 10 July 2007....

      • Podocarpus sprucei
        Podocarpus sprucei
        Podocarpus sprucei is a species of conifer in the Podocarpaceae family.It is found largely in Ecuador and Peru.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus transiens
        Podocarpus transiens
        Podocarpus transiens is a species of conifer in the Podocarpaceae family.It is found only in Brazil.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

    • section Pratensis (southeast Mexico
      Mexico
      The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

       to Guyana
      Guyana
      Guyana , officially the Co-operative Republic of Guyana, previously the colony of British Guiana, is a sovereign state on the northern coast of South America that is culturally part of the Anglophone Caribbean. Guyana was a former colony of the Dutch and of the British...

       and Peru
      Peru
      Peru , officially the Republic of Peru , is a country in western South America. It is bordered on the north by Ecuador and Colombia, on the east by Brazil, on the southeast by Bolivia, on the south by Chile, and on the west by the Pacific Ocean....

      )
      • Podocarpus oleifolius
        Podocarpus oleifolius
        Podocarpus oleifolius is a species of conifer in the Podocarpaceae family.It is found in Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Panama, Peru, and Venezuela.-References:...

      • Podocarpus pendulifolius
        Podocarpus pendulifolius
        Podocarpus pendulifolius is a species of conifer in the Podocarpaceae family.It is found only in Venezuela.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus tepuiensis
        Podocarpus tepuiensis
        Podocarpus tepuiensis is a species of conifer in the Podocarpaceae family.It is found only in Venezuela.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

    • section Lanceolatis (southern Mexico, Puerto Rico
      Puerto Rico
      Puerto Rico , officially the Commonwealth of Puerto Rico , is an unincorporated territory of the United States, located in the northeastern Caribbean, east of the Dominican Republic and west of both the United States Virgin Islands and the British Virgin Islands.Puerto Rico comprises an...

      , Lesser Antilles
      Lesser Antilles
      The Lesser Antilles are a long, partly volcanic island arc in the Western Hemisphere. Most of its islands form the eastern boundary of the Caribbean Sea with the Atlantic Ocean, with the remainder located in the southern Caribbean just north of South America...

      , Venezuela
      Venezuela
      Venezuela , officially called the Bolivarian Republic of Venezuela , is a tropical country on the northern coast of South America. It borders Colombia to the west, Guyana to the east, and Brazil to the south...

       to highland Bolivia
      Bolivia
      Bolivia officially known as Plurinational State of Bolivia , is a landlocked country in central South America. It is the poorest country in South America...

      )
      • Podocarpus coriaceus
        Podocarpus coriaceus
        Podocarpus coriaceus is a species of conifer in the Podocarpaceae family.It is found in the Dominican Republic, Guadeloupe, Martinique, Montserrat, Puerto Rico, and Saint Kitts and Nevis.-References:...

      • Podocarpus matudai
      • Podocarpus rusbyi
        Podocarpus rusbyi
        Podocarpus rusbyi is a species of conifer in the Podocarpaceae family.It is found only in Bolivia.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus salicifolius
        Podocarpus salicifolius
        Podocarpus salicifolius is a species of conifer in the Podocarpaceae family.It is found in Bolivia, Brazil, and Venezuela.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus steyermarkii
        Podocarpus steyermarkii
        Podocarpus steyermarkii is a species of conifer in the Podocarpaceae family.It is found only in Venezuela.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

    • section Pumilis (southern Caribbean islands and Guyana
      Guyana
      Guyana , officially the Co-operative Republic of Guyana, previously the colony of British Guiana, is a sovereign state on the northern coast of South America that is culturally part of the Anglophone Caribbean. Guyana was a former colony of the Dutch and of the British...

       highlands)
      • Podocarpus angustifolius
        Podocarpus angustifolius
        Podocarpus angustifolius is a species of conifer in the Podocarpaceae family.It is found only in Cuba.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus aristulatus
        Podocarpus aristulatus
        Podocarpus aristulatus is a small to medium-sized evergreen tree in the conifer family Podocarpaceae. It is found on the Caribbean islands of Hispaniola and Cuba.-Description:...

      • Podocarpus buchholzii
        Podocarpus buchholzii
        Podocarpus buchholzii is a species of conifer in the Podocarpaceae family.It is found only in Venezuela.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus roraimae
        Podocarpus roraimae
        Podocarpus roraimae is a species of conifer in the Podocarpaceae family.It is found only in Venezuela.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus urbanii
        Podocarpus urbanii
        Podocarpus urbanii is a species of conifer in the Podocarpaceae family.It is found only in Jamaica.-Source:* Bellingham, P. 1998. . Downloaded on 10 July 2007....

    • section Nemoralis (central and northern South America
      South America
      South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

      , south to Bolivia)
      • Podocarpus brasiliensis
        Podocarpus brasiliensis
        Podocarpus brasiliensis is a species of conifer in the Podocarpaceae family.It is found in Brazil and Venezuela.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus celatus
        Podocarpus celatus
        Podocarpus celatus is a species of conifer in the Podocarpaceae family.It is found in Bolivia, Peru, and Venezuela.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus guatemalensis
        Podocarpus guatemalensis
        Podocarpus guatemalensis, the Cipresillo or Pinillo, is a species of conifer in the Podocarpaceae family.-Distribution:Podocarpus guatemalensis is shrub and small tree found in namesake Guatemala, and also from southern Mexico through Belize, Costa Rica, El Salvador, Honduras, and Panama in Central...

      • Podocarpus magnifolius
        Podocarpus magnifolius
        Podocarpus magnifolius is a species of conifer in the Podocarpaceae family.It is found in Bolivia, Colombia, Peru, and Venezuela.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus purdieanus
        Podocarpus purdieanus
        Podocarpus purdieanus is a species of conifer in the Podocarpaceae family.It is found only in Jamaica.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus trinitensis
        Podocarpus trinitensis
        Podocarpus trinitensis is a species of conifer in the Podocarpaceae family. It is endemic to Trinidad and Tobago, where it has only been recorded from the island of Trinidad...


  • Subgenus Foliolatus
    • section Foliolatus (Nepal
      Nepal
      Nepal , officially the Federal Democratic Republic of Nepal, is a landlocked sovereign state located in South Asia. It is located in the Himalayas and bordered to the north by the People's Republic of China, and to the south, east, and west by the Republic of India...

       to Sumatra
      Sumatra
      Sumatra is an island in western Indonesia, westernmost of the Sunda Islands. It is the largest island entirely in Indonesia , and the sixth largest island in the world at 473,481 km2 with a population of 50,365,538...

      , Philippines
      Philippines
      The Philippines , officially known as the Republic of the Philippines , is a country in Southeast Asia in the western Pacific Ocean. To its north across the Luzon Strait lies Taiwan. West across the South China Sea sits Vietnam...

      , and New Guinea
      New Guinea
      New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...

       to Tonga
      Tonga
      Tonga, officially the Kingdom of Tonga , is a state and an archipelago in the South Pacific Ocean, comprising 176 islands scattered over of ocean in the South Pacific...

      )
      • Podocarpus archboldii
        Podocarpus archboldii
        Podocarpus archboldii is a species of conifer in the Podocarpaceae family.It is found in Indonesia and Papua New Guinea.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus beecherae
      • Podocarpus borneensis
        Podocarpus borneensis
        Podocarpus borneensis is a species of conifer in the Podocarpaceae family.It is found in Indonesia and Malaysia.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus deflexus
        Podocarpus deflexus
        Podocarpus deflexus is a species of conifer in the Podocarpaceae family.It is found in Indonesia and Malaysia.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus insularis
        Podocarpus insularis
        Podocarpus insularis is a species of conifer in the Podocarpaceae family.It is found in Indonesia, Papua New Guinea, Solomon Islands, and Vanuatu.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus levis
        Podocarpus levis
        Podocarpus levis is a species of conifer in the Podocarpaceae family.It is found in Brunei, Indonesia, and Malaysia.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus neriifolius
        Podocarpus neriifolius
        Podocarpus neriifolius is a species of conifer in the Podocarpaceae family.It is found in Brunei, Cambodia, China, Fiji, India, Indonesia, Laos, Malaysia, Myanmar, Nepal, Papua New Guinea, the Philippines, Solomon Islands, Thailand, and Vietnam....

      • Podocarpus novae-caledoniae
        Podocarpus novae-caledoniae
        Podocarpus novae-caledoniae is a species of conifer in the Podocarpaceae family.It is found only in New Caledonia.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus pallidus
        Podocarpus pallidus
        Podocarpus pallidus is a species of conifer in the Podocarpaceae family.It is found only in, and so endemic to Tonga in the South Pacific.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus rubens
        Podocarpus rubens
        Podocarpus rubens is a species of conifer in the Podocarpaceae family.It is found in Indonesia, Malaysia, Papua New Guinea, and Timor-Leste.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus spathoides
        Podocarpus spathoides
        Podocarpus spathoides is a species of conifer in the Podocarpaceae family.It is found in Indonesia, Malaysia, Papua New Guinea, and Solomon Islands.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

    • section Acuminatus (northern Queensland, New Guinea, New Britain
      New Britain
      New Britain, or Niu Briten, is the largest island in the Bismarck Archipelago of Papua New Guinea. It is separated from the island of New Guinea by the Dampier and Vitiaz Straits and from New Ireland by St. George's Channel...

      , Borneo
      Borneo
      Borneo is the third largest island in the world and is located north of Java Island, Indonesia, at the geographic centre of Maritime Southeast Asia....

      )
      • Podocarpus dispermus
        Podocarpus dispermus
        Podocarpus dispermus is a species of conifer in the Podocarpaceae family.It is found only in Queensland, Australia.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus ledermannii
        Podocarpus ledermannii
        Podocarpus ledermannii is a species of conifer in the Podocarpaceae family.It is found in Indonesia and Papua New Guinea.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus micropedunculatis
    • section Globulus (Taiwan
      Taiwan
      Taiwan , also known, especially in the past, as Formosa , is the largest island of the same-named island group of East Asia in the western Pacific Ocean and located off the southeastern coast of mainland China. The island forms over 99% of the current territory of the Republic of China following...

       to Vietnam
      Vietnam
      Vietnam – sometimes spelled Viet Nam , officially the Socialist Republic of Vietnam – is the easternmost country on the Indochina Peninsula in Southeast Asia. It is bordered by China to the north, Laos to the northwest, Cambodia to the southwest, and the South China Sea –...

      , Sumatra and Borneo, and New Caledonia)
      • Podocarpus annamiensis
        Podocarpus annamiensis
        Podocarpus annamiensis is a species of conifer in the Podocarpaceae family.It is found in China, Myanmar, and Vietnam.It is threatened by habitat loss.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus globulus
        Podocarpus globulus
        Podocarpus globulus is a species of conifer in the Podocarpaceae family.It is found in Indonesia and Malaysia.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus lucienii
        Podocarpus lucienii
        Podocarpus lucienii is a species of conifer in the Podocarpaceae family.It is found only in New Caledonia.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus nakai
      • Podocarpus sylvestris
        Podocarpus sylvestris
        Podocarpus sylvestris is a species of conifer in the Podocarpaceae family.It is found only in New Caledonia.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus teysmannii
        Podocarpus teysmannii
        Podocarpus teysmannii is a species of conifer in the Podocarpaceae family.It is found in Indonesia and Malaysia.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

    • section Longifoliolatus (Sumatra and Borneo, East to Fiji
      Fiji
      Fiji , officially the Republic of Fiji , is an island nation in Melanesia in the South Pacific Ocean about northeast of New Zealand's North Island...

      )
      • Podocarpus atjehensis
        Podocarpus atjehensis
        Podocarpus atjehensis is a species of conifer in the Podocarpaceae family.It is found in Indonesia and Papua New Guinea.-External Links:* on the Missouri Botanical Garden* on The Plant List...

      • Podocarpus bracteatus
        Podocarpus bracteatus
        Podocarpus bracteatus is a species of conifer in the Podocarpaceae family.It is found only in Indonesia.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus confertus
        Podocarpus confertus
        Podocarpus confertus is a species of conifer in the Podocarpaceae family.It is found in Brunei, Indonesia, and Malaysia.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus decumbens
        Podocarpus decumbens
        Podocarpus decumbens is a species of conifer in the Podocarpaceae family.It is found only in New Caledonia.It is threatened by habitat loss.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus degeneri
        Podocarpus degeneri
        Podocarpus degeneri is a species of conifer in the Podocarpaceae family.It is found only in Fiji.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus gibbsii
        Podocarpus gibbsii
        Podocarpus gibbsii is a species of conifer in the Podocarpaceae family.It is found only in Malaysia.It is threatened by habitat loss.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus longifoliolatus
        Podocarpus longifoliolatus
        Podocarpus longifoliolatus is a species of conifer in the Podocarpaceae family.It is found only in New Caledonia.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus polyspermus
        Podocarpus polyspermus
        Podocarpus polyspermus is a species of conifer in the Podocarpaceae family.It is found only in New Caledonia.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus pseudobracteatus
        Podocarpus pseudobracteatus
        Podocarpus pseudobracteatus is a species of conifer in the Podocarpaceae family.It is found in Indonesia and Papua New Guinea.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus salomoniensis
        Podocarpus salomoniensis
        Podocarpus salomoniensis is a species of conifer in the Podocarpaceae family.It is found only in Solomon Islands.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

    • section Gracilis (southern China
      China
      Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

      , across Malesia
      Malesia
      Malesia is a biogeographical region straddling the boundaries of the Indomalaya ecozone and Australasia ecozone, and also a phytogeographical floristic region in the Paleotropical Kingdom.-Floristic province:...

       to Fiji)
      • Podocarpus affinis
        Podocarpus affinis
        Podocarpus affinis is a species of conifer in the Podocarpaceae family.It is found only in Fiji.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus glaucus
        Podocarpus glaucus
        Podocarpus glaucus is a species of conifer in the Podocarpaceae family.It is found in Brunei, Indonesia, Malaysia, Papua New Guinea, the Philippines, and Solomon Islands.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus lophatus
        Podocarpus lophatus
        Podocarpus lophatus is a species of conifer in the Podocarpaceae family.It is found only in the Philippines.It is threatened by habitat loss.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus pilgeri
        Podocarpus pilgeri
        Podocarpus pilgeri is a species of conifer in the Podocarpaceae family.It is found in Cambodia and Thailand.-References:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus rotundus
        Podocarpus rotundus
        Podocarpus rotundus is a species of conifer in the Podocarpaceae family.It is found in Indonesia and the Philippines.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

    • section Macrostachyus (Southeast Asia
      Southeast Asia
      Southeast Asia, South-East Asia, South East Asia or Southeastern Asia is a subregion of Asia, consisting of the countries that are geographically south of China, east of India, west of New Guinea and north of Australia. The region lies on the intersection of geological plates, with heavy seismic...

       to New Guinea)
      • Podocarpus brassii
        Podocarpus brassii
        Podocarpus brassii is a species of conifer in the Podocarpaceae family.It is found in Indonesia and Papua New Guinea.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus brevifolius
        Podocarpus brevifolius
        Podocarpus brevifolius is a species of conifer in the Podocarpaceae family.It is found only in Malaysia.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus costalis
        Podocarpus costalis
        Podocarpus costalis is a species of conifer in the Podocarpaceae family.It is found in the Philippines and Taiwan.It is threatened by habitat loss.-References:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus crassigemmis
        Podocarpus crassigemmis
        Podocarpus crassigemmis is a species of conifer in the Podocarpaceae family.It is found in Indonesia and Papua New Guinea.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus tixieri
    • section Rumphius (Hainan
      Hainan
      Hainan is the smallest province of the People's Republic of China . Although the province comprises some two hundred islands scattered among three archipelagos off the southern coast, of its land mass is Hainan Island , from which the province takes its name...

      , south through Malesia to northern Queensland)
      • Podocarpus grayii
      • Podocarpus laubenfelsii
        Podocarpus laubenfelsii
        Podocarpus laubenfelsii is a species of conifer in the Podocarpaceae family.It is found only in Indonesia.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus rumphii
        Podocarpus rumphii
        Podocarpus rumphii is a species of conifer in the Podocarpaceae family.It is found in China, Indonesia, Malaysia, Papua New Guinea, and the Philippines.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

    • section Polystachyus (southern China and Japan
      Japan
      Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

      , through Malaya
      Malay Peninsula
      The Malay Peninsula or Thai-Malay Peninsula is a peninsula in Southeast Asia. The land mass runs approximately north-south and, at its terminus, is the southern-most point of the Asian mainland...

       to New Guinea and northeast Australia)
      • Podocarpus chinensis
        Podocarpus chinensis
        Podocarpus chinensis is a species of conifer in the Podocarpaceae family.It is found in China, Japan, and Myanmar.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus chingianus
      • Podocarpus elatus
        Podocarpus elatus
        Podocarpus elatus, known as the Plum Pine, or the Brown Pine is a species of Podocarpus endemic to the east coast of Australia, in eastern New South Wales and eastern Queensland....

      • Podocarpus fasciculus
        Podocarpus fasciculus
        Podocarpus fasciculus is a species of conifer in the Podocarpaceae family.It is found in Japan and Taiwan.It is threatened by habitat loss.-Source:* Conifer Specialist Group 2000. . Downloaded on 10 July 2007....

      • Podocarpus macrocarpus
        Podocarpus macrocarpus
        Podocarpus macrocarpus is a species of conifer in the Podocarpaceae family.It is found only in the Philippines.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus macrophyllus
        Podocarpus macrophyllus
        Podocarpus macrophyllus is a conifer in the genus Podocarpus, family Podocarpaceae. It is the northernmost species of the genus, native to southern Japan and China...

      • Podocarpus polystachyus
        Podocarpus polystachyus
        Podocarpus polystachyus is a species of conifer in the Podocarpaceae family.It is found in Brunei, Indonesia, Malaysia, Papua New Guinea, and the Philippines.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus ridleyi
        Podocarpus ridleyi
        Podocarpus ridleyi is a species of conifer in the Podocarpaceae family.It is found only in Malaysia.-References:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

      • Podocarpus subtropicalis
        Podocarpus subtropicalis
        Podocarpus subtropicalis is a species of conifer in the Podocarpaceae family.It is found only in China.-Source:* Conifer Specialist Group 1998. . Downloaded on 10 July 2007....

    • section Spinulosus (Southeast and Southwest coasts of Australia)
      • Podocarpus drouynianus
        Podocarpus drouynianus
        Podocarpus drouynianus is a species of podocarp native to the relatively high rainfall southwestern corner of Western Australia, where it is known by the name Wild Plum, although it is not a true plum. It grows around creeks in sandy or gravelly soil. It is usually a shrub, not often forming a...

      • Podocarpus spinulosus
        Podocarpus spinulosus
        Podocarpus spinulosus, the Dwarf Plum Pine or Spiny-leaf Podocarp, is a species of podocarp native to the warm-temperate coastal regions of New South Wales and southern Queensland. It is generally an understorey shrub, rarely growing more than 2 m tall.It was first described by James Edward...


Uses

Several species of Podocarpus are grown as garden trees, or trained into hedges, espaliers, or screens. Common garden species used for their attractive deep green foliage and neat habits include P. macrophyllus, known by its Japanese name Kusamaki, or occasionally as "buddhist pine" or "fern pine", P. salignus from Chile, and for a small shrub with attractive red "berries", P. nivalis. Some members of the genera Nageia
Nageia
Nageia is a genus of conifers belonging to the podocarp family Podocarpaceae. Nageia includes evergreen shrubs and trees, from one to 54 meters in height. A 2009 treatment of the genus recognized five species. Some authors consider Nageia formosensis to be a separate species from Nageia nagi,...

, Prumnopitys
Prumnopitys
Prumnopitys is a genus of conifers belonging to the podocarp family Podocarpaceae. The eight recognised species of Prumnopitys are densely-branched, dioecious evergreen trees up to 40 metres in height. The leaves are similar to those of the yew, strap-shaped, 1-4 cm long and 2-3 mm broad, with a...

and Afrocarpus
Afrocarpus
Afrocarpus is a genus of conifers belonging to the podocarp family Podocarpaceae. Afrocarpus was designated a genus in 1989, when several species formerly classified in Podocarpus and Nageia were reclassified. Two to six species are recognized.As the name intimates, Afrocarpus is native to Africa...

are also still sold mislabeled as Podocarpus. The red, purple or bluish fleshy fruit of most species of Podocarpus are edible, raw or cooked into jams or pies, and they have a mucilaginous texture with a slightly sweet flavor. However, the fruit are slightly toxic and should therefore be eaten sparingly, especially when eaten raw. The timber of P. falcatus is used for floorboards, beams and furniture.

Some species of Podocarpus are used traditionally in their native areas for the treatment of fevers, asthma, coughs, cholera, chest complaints, arthritis, rheumatism, venereal diseases and distemper in dogs.
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