Pupilla muscorum
Encyclopedia
Pupilla muscorum is a species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of minute air-breathing land snail
Land snail
A land snail is any of the many species of snail that live on land, as opposed to those that live in salt water and fresh water. Land snails are terrestrial gastropod mollusks that have shells, It is not always an easy matter to say which species are terrestrial, because some are more or less...

, a terrestrial
Terrestrial animal
Terrestrial animals are animals that live predominantly or entirely on land , as compared with aquatic animals, which live predominantly or entirely in the water , or amphibians, which rely on a combination of aquatic and terrestrial habitats...

 pulmonate gastropod mollusk or micromollusk
Micromollusk
A micromollusk is a descriptive term for a shelled mollusk which is extremely small, even at full adult size. The word is usually, but not exclusively, applied to marine mollusks, although in addition, numerous species of land snails and freshwater mollusks also reach adult size at very small...

 in the family Pupillidae
Pupillidae
Pupillidae is a family of mostly minute, air-breathing, land snails, terrestrial pulmonate gastropod mollusks or micromollusks in the clade Stylommatophora....

.

Taxonomy

A variety Pupilla muscorum var. pratensis Clessin, 1871 considered as ecophenotype of Pupilla muscorum was elevated to its species level Pupilla pratensis
Pupilla pratensis
Pupilla pratensis is a species of minute air-breathing land snail, a terrestrial pulmonate gastropod mollusk or micromollusk in the family Pupillidae. This taxon was elevated to species level in 2009.- Taxonomy :...

in 2009.

Description

The shell
Gastropod shell
The gastropod shell is a shell which is part of the body of a gastropod or snail, one kind of mollusc. The gastropod shell is an external skeleton or exoskeleton, which serves not only for muscle attachment, but also for protection from predators and from mechanical damage...

 is usually light brown, varies from reddish brown to horny grey, weakly striated or almost smooth, 5-6.5 weakly convex whorls
Whorl (mollusc)
A whorl is a single, complete 360° revolution or turn in the spiral growth of a mollusc shell. A spiral configuration of the shell is found in of numerous gastropods, but it is also found in shelled cephalopods including Nautilus, Spirula and the large extinct subclass of cephalopods known as the...

, suture not very deep, aperture
Aperture (mollusc)
The aperture is an opening in certain kinds of mollusc shells: it is the main opening of the shell, where part of the body of the animal emerges for locomotion, feeding, etc....

 usually with well-developed lip, cervical callus strongly developed, like a dam, parietal tooth usually present, palatal tooth sometimes too.

Pupilla muscorum differs from Pupilla pratensis with which it lives sympatrically, in its thicker, smaller and more slender shell, lighter and more variable colour and stronger apertural lip.

The animal of Pupilla muscorum is small, elliptical, dark with lighter sides and foot, upper tentacles not very long, lower tentacles very short.

The height of the shell is 3.0-4.0 mm. The width of the shell is 1.65-1.75 mm (shell diameter does not vary much).

Distribution

This species of land snail occurs in the Northern Hemisphere including almost all of Europe. It is recorded as present in countries and islands including:

Ecology

Pupilla muscorum lives on dry meadows, sand dunes, open and sunny habitats. Calciphile. In Portugal found under stones, dead leaves and in mosses. In Britain frequent in sheep-grazed calcareous grasslands. In the Alps in up to 2400 m, in Bulgaria 1200 m.

Ovoviviparous, the species is able to hibernate with eggs and can release eggs with partly grown embryos in favourable seasons.

External links

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