Purussaurus
Encyclopedia
Purussaurus is an extinct genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of giant caiman
Caiman
Caimans are alligatorid crocodylians within the subfamily Caimaninae. The group is one of two subfamilies of the family Alligatoridae, the other being alligators. Caimans inhabit Central and South America. They are relatively small crocodilians, with most species reaching lengths of only a few...

 that lived in South America
South America
South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

 during the Miocene
Miocene
The Miocene is a geological epoch of the Neogene Period and extends from about . The Miocene was named by Sir Charles Lyell. Its name comes from the Greek words and and means "less recent" because it has 18% fewer modern sea invertebrates than the Pliocene. The Miocene follows the Oligocene...

 epoch, 8 million years ago. It is known from skull material found in the Brazilian, Colombian and Peru
Peru
Peru , officially the Republic of Peru , is a country in western South America. It is bordered on the north by Ecuador and Colombia, on the east by Brazil, on the southeast by Bolivia, on the south by Chile, and on the west by the Pacific Ocean....

vian Amazonia, and northern Venezuela. The estimated skull length for one large individual of the type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

 P. brasiliensis is 1400 millimetres (55.1 in). Paleontologists estimate that P. brasiliensis may have measured around 11 to 13 m (36.1 to 42.7 ) in length, which means that Purussaurus is one of the largest crocodilians known to have ever existed. Two other extinct crocodilians, Sarcosuchus
Sarcosuchus
Sarcosuchus is an extinct genus of crocodyliform and distant relative of the crocodile that lived 112 million years ago. It dates from the early Cretaceous Period of what is now Africa and is one of the largest giant crocodile-like reptiles that ever lived...

and Deinosuchus
Deinosuchus
Deinosuchus is an extinct genus related to the alligator that lived 73 to 80 Ma , during the late Cretaceous period. The name translates as "terrible crocodile" and is derived from the Greek deinos , "terrible", and soukhos , "crocodile"...

, have similar proportions, but both are geologically much older, dating from the Early and Late Cretaceous
Cretaceous
The Cretaceous , derived from the Latin "creta" , usually abbreviated K for its German translation Kreide , is a geologic period and system from circa to million years ago. In the geologic timescale, the Cretaceous follows the Jurassic period and is followed by the Paleogene period of the...

, respectively, and another from the Miocene of India
India
India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

, Rhamphosuchus
Rhamphosuchus
Rhamphosuchus is an extinct relative of the modern false gharial. It inhabited what is now the Indian sub-continent in the Miocene...

, is estimated to be slightly smaller, though assumed to have been proportioned like a gharial. During the summer of 2005, a Franco-Peruvian expedition (the Fitzcarrald expedition) found new fossil
Fossil
Fossils are the preserved remains or traces of animals , plants, and other organisms from the remote past...

s of Purussaurus in the Peruvian Amazon
Peruvian Amazon
The Peruvian Amazon is the area of the Amazon jungle included in the territory of Peru, from the east of the Andes to borders with Ecuador, Colombia, Brazil and Bolivia. This region comprises 60% of the country and is marked by a large degree of biodiversity...

 (600 km from Lima
Lima
Lima is the capital and the largest city of Peru. It is located in the valleys of the Chillón, Rímac and Lurín rivers, in the central part of the country, on a desert coast overlooking the Pacific Ocean. Together with the seaport of Callao, it forms a contiguous urban area known as the Lima...

).

Paleoecology

Brazilian P. brasiliensis is associated with shark
Shark
Sharks are a type of fish with a full cartilaginous skeleton and a highly streamlined body. The earliest known sharks date from more than 420 million years ago....

s, rays
Batoidea
Batoidea is a superorder of cartilaginous fish commonly known as rays and skates, containing more than 500 described species in thirteen families...

, freshwater teleosts, lungfish
Lungfish
Lungfish are freshwater fish belonging to the Subclass Dipnoi. Lungfish are best known for retaining characteristics primitive within the Osteichthyes, including the ability to breathe air, and structures primitive within Sarcopterygii, including the presence of lobed fins with a well-developed...

, turtles including Stupendemys
Stupendemys
Stupendemys is a prehistoric genus of freshwater turtle. Its fossils have been found in northern South America, in rocks dating from the late Miocene to the very start of the Pliocene, about 6 to 5 million years ago....

, crocodilians including Charactosuchus
Charactosuchus
Charactosuchus is an extinct genus of crocodilian. It was assigned to the family Crocodylidae in 1988. Specimens have been found from Colombia, Brazil, Jamaica, and possibly Florida and South Carolina...

, Gryposuchus
Gryposuchus
Gryposuchus is an extinct genus of gavialoid crocodilian. It is the type genus of the subfamily Gryposuchinae. Fossils have been found from Argentina, Colombia, Venezuela, Brazil and the Peruvian Amazon. The genus existed during the early and middle Miocene epoch. One recently described species, G...

, and Mourasuchus
Mourasuchus
Mourasuchus is an extinct genus of giant crocodilian from the Miocene of South America. It was similar in length and weight to Rhamphosuchus...

, the bird Anhinga
Anhinga
The Anhinga , sometimes called Snakebird, Darter, American Darter, or Water Turkey, is a water bird of the warmer parts of the Americas. The word "anhinga" comes from the Brazilian Tupi language and means devil bird or snake bird.It is a cormorant-like bird with an average body length of , a...

, and mammals including sloth
Sloth
Sloths are the six species of medium-sized mammals belonging to the families Megalonychidae and Bradypodidae , part of the order Pilosa and therefore related to armadillos and anteaters, which sport a similar set of specialized claws.They are arboreal residents of the jungles of Central and South...

s, bats, rodents, the primate Stirtonia, and river dolphins. River, floodplain, and lake environments were present. Marine and freshwater fish, turtles, crocodilians, and terrestrial and aquatic mammals are associated with Venezuelan P. mirandai. Its environment is described as tropical and coastal.

Sources

  • Rodrigues, J.B. 1892. "Les reptiles fossils de la Vallée de L’Amazone". Vellosia, Contribuições do Museu Botânico do Amazonas 2:41-60.
  • Langston, W. 1965. "Fossil crocodilians from Colombia and the Cenozoic history of the Crocodilia in South America". University of California Publications in Geological Sciences 52:1-169.

External links

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