Rhinichthys
Encyclopedia
Rhinichthys, the riffle dace
s, is a genus
of freshwater
fish
in the carp family
(Cyprinidae) of the order
Cypriniformes
. The type species
is Rhinichthys atratulus, the blacknose dace. Rhinichthys species range throughout North America
.
The genus contains eight living species, one of which (the loach minnow) is considered Vulnerable. It also includes the extinct Las Vegas dace, which was only first described in 1984 and had disappeared by 1986. The cheat minnow
(Pararhinichthys bowersi), a natural hybrid of the longnose dace (R. cataractae) and the river chub (Nocomis micropogon), was formerly placed in this genus, but is now valid under Pararhinichthys.
The riffle daces are a basal
lineage in an insufficiently resolved clade
of American Leuciscinae
. Such a group had been proposed on anatomical evidence, and was verified using mtDNA 12S rRNA sequence
s.(Simons & Mayden 1997)
Dace
A dace is any of a number of species of small fish. The unmodified name is usually a reference to the Common Dace . This, like most fishes called "daces", belongs to the family Cyprinidae, mostly in subfamily Leuciscinae....
s, is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...
of freshwater
Freshwater
Fresh water is naturally occurring water on the Earth's surface in ice sheets, ice caps, glaciers, bogs, ponds, lakes, rivers and streams, and underground as groundwater in aquifers and underground streams. Fresh water is generally characterized by having low concentrations of dissolved salts and...
fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...
in the carp family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...
(Cyprinidae) of the order
Order (biology)
In scientific classification used in biology, the order is# a taxonomic rank used in the classification of organisms. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, and species, with order fitting in between class and family...
Cypriniformes
Cypriniformes
The Cypriniformes are an order of ray-finned fish, including the carps, minnows, loaches and relatives. This order contains 5-6 families, over 320 genera, and more than 3,250 species, with new species being described every few months or so, and new genera being recognized regularly...
. The type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...
is Rhinichthys atratulus, the blacknose dace. Rhinichthys species range throughout North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...
.
The genus contains eight living species, one of which (the loach minnow) is considered Vulnerable. It also includes the extinct Las Vegas dace, which was only first described in 1984 and had disappeared by 1986. The cheat minnow
Cheat minnow
The cheat minnow is a demersal, freshwater fish endemic to the United States, most commonly found in the Ohio River basin...
(Pararhinichthys bowersi), a natural hybrid of the longnose dace (R. cataractae) and the river chub (Nocomis micropogon), was formerly placed in this genus, but is now valid under Pararhinichthys.
The riffle daces are a basal
Basal (phylogenetics)
In phylogenetics, a basal clade is the earliest clade to branch in a larger clade; it appears at the base of a cladogram.A basal group forms an outgroup to the rest of the clade, such as in the following example:...
lineage in an insufficiently resolved clade
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...
of American Leuciscinae
Leuciscinae
Leuciscinae, commonly known as true minnows, are a subfamily of small fish belonging to the family Cyprinidae.However, the term minnow is also used as an unspecific term for tiny freshwater and saltwater fish, especially those used as fishing bait....
. Such a group had been proposed on anatomical evidence, and was verified using mtDNA 12S rRNA sequence
DNA sequence
The sequence or primary structure of a nucleic acid is the composition of atoms that make up the nucleic acid and the chemical bonds that bond those atoms. Because nucleic acids, such as DNA and RNA, are unbranched polymers, this specification is equivalent to specifying the sequence of...
s.(Simons & Mayden 1997)
Species
- Rhinichthys atratulus (HermannJohann HermannJohann, or Jean, Hermann was a French physician and naturalist. He was professor of medicine at the University of Strasbourg. He was the author of Tabula affinitatum animalium and Observationes zoologicae quibus novae complures, published posthumously in 1804...
, 1804) (Blacknose dace) - Rhinichthys cataractae (ValenciennesAchille ValenciennesAchille Valenciennes was a French zoologist.Valenciennes was born in Paris, and studied under Georges Cuvier. Valenciennes' study of parasitic worms in humans made an important contribution to the study of parasitology...
, 1842) (Longnose dace) - Rhinichthys cobitis (GirardCharles Frédéric GirardCharles Frédéric Girard was a French biologist specializing in ichthyology and herpetology.Born in Mulhouse, France, he studied at the College of Neuchâtel, Switzerland as a student of Louis Agassiz. In 1847, he accompanied Agassiz as his assistant to Harvard...
, 1856) (Loach minnow) - †Rhinichthys deaconi R. R. MillerRobert Rush MillerRobert Rush Miller "was an important figure in American ichthyology and conservation from 1940 to the 1990s."...
, 1984 (Las Vegas Dace) - Rhinichthys evermanni SnyderJohn Otterbein SnyderJohn Otterbein Snyder was an American zoologist.As a student he met David Starr Jordan who inspired him to enter zoology. He eventually became a zoology instructor at Stanford University and served there from 1899 until 1943. He went on several major collecting expeditions aboard the USS Albatross...
, 1908 (Umpqua dace) - Rhinichthys falcatus (C. H. EigenmannCarl H. EigenmannCarl H. Eigenmann was an ichthyologist who, along with his wife Rosa Smith Eigenmann, described many of the fishes of North America and South America for the first time....
& R. S. EigenmannRosa Smith EigenmannRosa Smith Eigenmann was the first notable female ichthyologist; first publishing in her own right, she later collaborated with her husband Carl H. Eigenmann, and some 150 species of fish are today credited "Eigenmann & Eigenmann" as a result.She was born in Monmouth, Illinois, the last of nine...
, 1893) (Leopard dace) - Rhinichthys obtusus AgassizLouis AgassizJean Louis Rodolphe Agassiz was a Swiss paleontologist, glaciologist, geologist and a prominent innovator in the study of the Earth's natural history. He grew up in Switzerland and became a professor of natural history at University of Neuchâtel...
, 1854 (Western blacknose dace) - Rhinichthys osculus (GirardCharles Frédéric GirardCharles Frédéric Girard was a French biologist specializing in ichthyology and herpetology.Born in Mulhouse, France, he studied at the College of Neuchâtel, Switzerland as a student of Louis Agassiz. In 1847, he accompanied Agassiz as his assistant to Harvard...
, 1856) (Speckled dace) - Rhinichthys umatilla (C. H. GilbertCharles Henry GilbertCharles Henry Gilbert was a pioneer ichthyologist and fishery biologist of particular significance to natural history of the western United States. He collected and studied fishes from Central America north to Alaska and described many new species...
& EvermannBarton Warren EvermannBarton Warren Evermann was an American ichthyologist. He was born in Monroe County, Iowa, and graduated from Indiana University in 1886. For 10 years, he served as teacher and superintendent of schools in Indiana and California. He was professor of biology at the Indiana State University in...
, 1894) (Umatilla dace)