Robert Brown was a Scottish botanist and palaeobotanist who made important contributions to botany largely through his pioneering use of the microscope. His contributions include the discovery of the cell nucleus

Cell nucleus

In cell biology, the nucleus is a membrane-enclosed organelle found in eukaryotic cells. It contains most of the cell's genetic material, organized as multiple long linear DNA molecules in complex with a large variety of proteins, such as histones, to form chromosomes. The genes within these...

 and cytoplasmic streaming

Cytoplasmic streaming

Cytoplasmic streaming is the directed flow of cytosol and organelles around large fungal and plant cells. This movement aids in the delivery of nutrients, metabolites, genetic information, and other materials to all parts of the cell...

; the first observation of Brownian motion

Brownian motion

Brownian motion or pedesis is the presumably random drifting of particles suspended in a fluid or the mathematical model used to describe such random movements, which is often called a particle theory.The mathematical model of Brownian motion has several real-world applications...

; early work on plant pollination

Pollination

Pollination is the process by which pollen is transferred in plants, thereby enabling fertilisation and sexual reproduction. Pollen grains transport the male gametes to where the female gamete are contained within the carpel; in gymnosperms the pollen is directly applied to the ovule itself...

 and fertilisation

Fertilisation

Fertilisation is the fusion of gametes to produce a new organism. In animals, the process involves the fusion of an ovum with a sperm, which eventually leads to the development of an embryo...

, including being the first to recognise the fundamental difference between gymnosperms and angiosperms; and some of the earliest studies in palynology

Palynology

Palynology is the science that studies contemporary and fossil palynomorphs, including pollen, spores, orbicules, dinoflagellate cysts, acritarchs, chitinozoans and scolecodonts, together with particulate organic matter and kerogen found in sedimentary rocks and sediments...

.