Lanner Falcon
Encyclopedia
The Lanner Falcon is a large bird of prey
Bird of prey
Birds of prey are birds that hunt for food primarily on the wing, using their keen senses, especially vision. They are defined as birds that primarily hunt vertebrates, including other birds. Their talons and beaks tend to be relatively large, powerful and adapted for tearing and/or piercing flesh....

 that breeds in Africa
Africa
Africa is the world's second largest and second most populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area...

, southeast Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

 and just into Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

. It is mainly resident, but some bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

s disperse
Bird migration
Bird migration is the regular seasonal journey undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather. Sometimes, journeys are not termed "true migration" because they are irregular or in only one direction...

 more widely after the breeding season.

Description

It is a large falcon
Falcon
A falcon is any species of raptor in the genus Falco. The genus contains 37 species, widely distributed throughout Europe, Asia, and North America....

, at 43–50 cm length with a wingspan of 95–105 cm. European Lanner Falcons (Falco biarmicus feldeggi, also called Feldegg's Falcon) have slate grey or brown-grey upperparts; most African subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

 are a paler blue grey above. The breast is streaked in northern birds, resembling greyish Saker Falcon
Saker Falcon
The Saker Falcon is a very large falcon. This species breeds from eastern Europe eastwards across Asia to Manchuria. It is mainly migratory except in the southernmost parts of its range, wintering in Ethiopia, the Arabian peninsula, northern Pakistan and western China...

s, but the Lanner has a reddish back to the head. Sexes are similar, but the browner young birds resemble Saker Falcons even more. However, Sakers have a lighter top of the head and less clear head-side patterns. The Lanner's call is a harsh "wray-e".

Taxonomy and systematics

The Lanner Falcon is a bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

 of open country and savanna
Savanna
A savanna, or savannah, is a grassland ecosystem characterized by the trees being sufficiently small or widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to support an unbroken herbaceous layer consisting primarily of C4 grasses.Some...

. It usually hunts by horizontal pursuit, rather than the Peregrine Falcon
Peregrine Falcon
The Peregrine Falcon , also known as the Peregrine, and historically as the Duck Hawk in North America, is a widespread bird of prey in the family Falconidae. A large, crow-sized falcon, it has a blue-gray back, barred white underparts, and a black head and "moustache"...

's stoop from a height, and takes mainly bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

 prey in flight. It lays 3-4 eggs
Bird egg
Bird eggs are laid by females and incubated for a time that varies according to the species; a single young hatches from each egg. Average clutch sizes range from one to about 17...

 on a cliff ledge nest
Bird nest
A bird nest is the spot in which a bird lays and incubates its eggs and raises its young. Although the term popularly refers to a specific structure made by the bird itself—such as the grassy cup nest of the American Robin or Eurasian Blackbird, or the elaborately woven hanging nest of the...

, or occasionally in an old stick nest in a tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

.

This is presumably the oldest living hierofalcon
Hierofalcon
The hierofalcons are four closely related species of falcon which make up the subgenus Hierofalco:* Lanner Falcon, Falco biarmicus* Laggar Falcon, Falco jugger* Saker Falcon, Falco cherrug* Gyrfalcon, Falco rusticolus...

 species. Support for this assumption comes mainly from biogeography
Biogeography
Biogeography is the study of the distribution of species , organisms, and ecosystems in space and through geological time. Organisms and biological communities vary in a highly regular fashion along geographic gradients of latitude, elevation, isolation and habitat area...

 agreeing better with the confusing pattern of DNA sequence
DNA sequence
The sequence or primary structure of a nucleic acid is the composition of atoms that make up the nucleic acid and the chemical bonds that bond those atoms. Because nucleic acids, such as DNA and RNA, are unbranched polymers, this specification is equivalent to specifying the sequence of...

 data in this case than in others. Nonetheless, there is rampant hybridization (see also Perilanner
Perilanner
Perilanner is a term used by falconers to describe a hybrid between a peregrine falcon and a lanner falcon. It is larger and faster than a lanner, but does not fly as far as a peregrine, and thus is less likely to fly far away and become lost. Usually the peregrine is the father and the lanner is...

) and incomplete lineage sorting which confounds the data to a massive extent; molecular studies with small sample size
Sample size
Sample size determination is the act of choosing the number of observations to include in a statistical sample. The sample size is an important feature of any empirical study in which the goal is to make inferences about a population from a sample...

s can simply not be expected to yield reliable conclusions in the entire hierofalcon group. In any case, the radiation of the entire living diversity of hierofalcons seems to have taken place in the Eemian interglacial
Eemian interglacial
The Eemian was an interglacial period which began about 130,000 years ago and ended about 114,000 years ago. It was the second-to-latest interglacial period of the current Ice Age, the most recent being the Holocene which extends to the present day. The prevailing Eemian climate is believed to...

 at the start of the Late Pleistocene
Late Pleistocene
The Late Pleistocene is a stage of the Pleistocene Epoch. The beginning of the stage is defined by the base of the Eemian interglacial phase before the final glacial episode of the Pleistocene 126,000 ± 5,000 years ago. The end of the stage is defined exactly at 10,000 Carbon-14 years BP...

, a mere 130,000-115,000 years ago; the Lanner Falcons would thus represent the lineage that became isolated in sub-Saharan Africa
Sub-Saharan Africa
Sub-Saharan Africa as a geographical term refers to the area of the African continent which lies south of the Sahara. A political definition of Sub-Saharan Africa, instead, covers all African countries which are fully or partially located south of the Sahara...

 at some time during the Riss glaciation (200,000 to 130,000 years ago) already.

They are bred in captivity
Captivity (animal)
Animals that live under human care are in captivity. Captivity can be used as a generalizing term to describe the keeping of either domesticated animals or wild animals. This may include for example farms, private homes and zoos...

 for falconry; hybrids with the Peregrine ("perilanner
Perilanner
Perilanner is a term used by falconers to describe a hybrid between a peregrine falcon and a lanner falcon. It is larger and faster than a lanner, but does not fly as far as a peregrine, and thus is less likely to fly far away and become lost. Usually the peregrine is the father and the lanner is...

s") are also often seen. Merret (1666) claimed that the "lanar" lived in Sherwood Forest
Sherwood Forest
Sherwood Forest is a Royal Forest in Nottinghamshire, England, that is famous through its historical association with the legend of Robin Hood. Continuously forested since the end of the Ice Age, Sherwood Forest National Nature Reserve today encompasses 423 hectares surrounding the village of...

 and the Forest of Dean
Forest of Dean
The Forest of Dean is a geographical, historical and cultural region in the western part of the county of Gloucestershire, England. The forest is a roughly triangular plateau bounded by the River Wye to the west and north, the River Severn to the south, and the City of Gloucester to the east.The...

 in England
England
England is a country that is part of the United Kingdom. It shares land borders with Scotland to the north and Wales to the west; the Irish Sea is to the north west, the Celtic Sea to the south west, with the North Sea to the east and the English Channel to the south separating it from continental...

; such populations would seem to derive from escaped hunting birds of the nobility.

In the wild Lanner Falcon numbers are somewhat declining in Europe, though the species remains relatively common in parts of Africa.

Jackdaw
Jackdaw
The Jackdaw , sometimes known as the Eurasian Jackdaw, European Jackdaw or Western Jackdaw, is a passerine bird in the crow family. Found across Europe, western Asia and North Africa, it is mostly sedentary, although northern and eastern populations migrate south in winter. Four subspecies are...

flocks are targets of coordinated hunting by pairs of Lanner Falcons, although larger flocks are more able to elude becoming prey.
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