Rachel Carson National Wildlife Refuge
Encyclopedia
The Rachel Carson National Wildlife Refuge is a 5400 acres (22 km²) National Wildlife Refuge
National Wildlife Refuge
National Wildlife Refuge is a designation for certain protected areas of the United States managed by the United States Fish and Wildlife Service. The National Wildlife Refuge System is the world's premiere system of public lands and waters set aside to conserve America's fish, wildlife and plants...

 made up of several parcels of land along 50 miles (80 km) of Maine
Maine
Maine is a state in the New England region of the northeastern United States, bordered by the Atlantic Ocean to the east and south, New Hampshire to the west, and the Canadian provinces of Quebec to the northwest and New Brunswick to the northeast. Maine is both the northernmost and easternmost...

's southern coast. Created in 1966, it is named for environmentalist
Environmentalist
An environmentalist broadly supports the goals of the environmental movement, "a political and ethical movement that seeks to improve and protect the quality of the natural environment through changes to environmentally harmful human activities"...

 and author Rachel Carson
Rachel Carson
Rachel Louise Carson was an American marine biologist and conservationist whose writings are credited with advancing the global environmental movement....

, whose book Silent Spring
Silent Spring
Silent Spring is a book written by Rachel Carson and published by Houghton Mifflin on 27 September 1962. The book is widely credited with helping launch the environmental movement....

raised public awareness of the effects of DDT
DDT
DDT is one of the most well-known synthetic insecticides. It is a chemical with a long, unique, and controversial history....

 on migratory
Bird migration
Bird migration is the regular seasonal journey undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather. Sometimes, journeys are not termed "true migration" because they are irregular or in only one direction...

 songbird
Songbird
A songbird is a bird belonging to the suborder Passeri of the perching birds . Another name that is sometimes seen as scientific or vernacular name is Oscines, from Latin oscen, "a songbird"...

s, and of other environmental issues.

The refuge's parcels include protected areas between Kittery
Kittery, Maine
Kittery is a town in York County, Maine, United States. The population was 9,543 at the 2000 census. Home to the Portsmouth Naval Shipyard on Seavey's Island, Kittery includes Badger's Island, the seaside district of Kittery Point, and part of the Isles of Shoals...

 and Cape Elizabeth
Cape Elizabeth, Maine
Cape Elizabeth is a town in Cumberland County, Maine, United States. The town is part of the Portland–South Portland–Biddeford, Maine metropolitan statistical area...

, including land in Wells
Wells, Maine
Wells is a town in York County, Maine, United States. Founded in 1643, it is the third-oldest town in Maine. The population was 9,400 at the 2000 census. Wells Beach is a popular summer destination.-History:...

, Kennebunk
Kennebunk, Maine
Kennebunk is a town in York County, Maine, United States. The population was 1,075 people at the 2000 census. Including Kennebunkport , the population totals 14,196 people...

, Kennebunkport
Kennebunkport, Maine
Kennebunkport is a town in York County, Maine, United States. The population was 3,720 at the 2000 census. It is part of the Portland–South Portland–Biddeford metropolitan statistical area....

, Biddeford
Biddeford, Maine
Biddeford is a town in York County, Maine, United States. It is the largest town in the county, and is the sixth-largest in the state. It is the most southerly incorporated town in the state and the principal commercial center of York County. The population was 21,277 at the 2010 census...

, Saco
Saco, Maine
Saco is a city in York County, Maine, United States. The population was 18,482 at the 2010 census. It is home to Ferry Beach State Park, Funtown Splashtown USA, Thornton Academy, as well as General Dynamics Armament Systems , a subsidiary of the defense contractor General Dynamics...

, and Scarborough.

The refuge protects 1167 acres (5 km²) of estuary salt marsh
Salt marsh
A salt marsh is an environment in the upper coastal intertidal zone between land and salt water or brackish water, it is dominated by dense stands of halophytic plants such as herbs, grasses, or low shrubs. These plants are terrestrial in origin and are essential to the stability of the salt marsh...

 and uplands
Upland and lowland (freshwater ecology)
In studies of the ecology of freshwater rivers, habitats are classified as upland and lowland.-Definitions:Upland habitats are cold, clear, rocky, fast flowing rivers in mountainous areas; lowland habitats are warm, slow flowing rivers found in relatively flat lowland areas, with water that is...

 that drain into the Webhannet River
Webhannet River
The Webhannet River is an river whose watershed is contained entirely within the town of Wells, Maine.The river has five tributaries, including three with official names: Pope’s Creek, Depot Brook, and Blacksmith Brook...

, or about one-ninth of the river's watershed.

The refuge's headquarters are on Route 9 in Wells.

The refuge protects various kinds of habitats, including barrier beach, dune
Dune
In physical geography, a dune is a hill of sand built by wind. Dunes occur in different forms and sizes, formed by interaction with the wind. Most kinds of dunes are longer on the windward side where the sand is pushed up the dune and have a shorter "slip face" in the lee of the wind...

, tidal estuary, salt marsh, and rocky coastline. The piping plover
Piping Plover
The Piping Plover is a small sand-colored, sparrow-sized shorebird that nests and feeds along coastal sand and gravel beaches in North America. The adult has yellow-orange legs, a black band across the forehead from eye to eye, and a black ring around the neck...

, an endangered species, nests on refuge land.

Wildlife and Habitat

The Rachel Carson National Wildlife Refuge was established to preserve ten important estuaries
Estuary
An estuary is a partly enclosed coastal body of water with one or more rivers or streams flowing into it, and with a free connection to the open sea....

 that are key points along migration routes
Bird migration
Bird migration is the regular seasonal journey undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather. Sometimes, journeys are not termed "true migration" because they are irregular or in only one direction...

 of waterfowl and other migratory birds. During harsh winters, the refuge's marshes provide vital food and cover for waterfowl and other migrating birds at a time when inland waters are frozen. The refuge also supports Piping Plover
Piping Plover
The Piping Plover is a small sand-colored, sparrow-sized shorebird that nests and feeds along coastal sand and gravel beaches in North America. The adult has yellow-orange legs, a black band across the forehead from eye to eye, and a black ring around the neck...

, Least Terns, Peregrine Falcon
Peregrine Falcon
The Peregrine Falcon , also known as the Peregrine, and historically as the Duck Hawk in North America, is a widespread bird of prey in the family Falconidae. A large, crow-sized falcon, it has a blue-gray back, barred white underparts, and a black head and "moustache"...

s, Bald Eagle
Bald Eagle
The Bald Eagle is a bird of prey found in North America. It is the national bird and symbol of the United States of America. This sea eagle has two known sub-species and forms a species pair with the White-tailed Eagle...

s and other state and federally protected species. Nesting success of plover and terns has benefitted through the increased habitat protection. In addition to anadromous fish, many commercially and recreationally important fin and shellfish rely on these coastal wetlands as critical nursery areas.

Refuge lands total over 5300 acres (21 km²) in eleven geographic units from Kittery
Kittery, Maine
Kittery is a town in York County, Maine, United States. The population was 9,543 at the 2000 census. Home to the Portsmouth Naval Shipyard on Seavey's Island, Kittery includes Badger's Island, the seaside district of Kittery Point, and part of the Isles of Shoals...

 to Cape Elizabeth, Maine
Cape Elizabeth, Maine
Cape Elizabeth is a town in Cumberland County, Maine, United States. The town is part of the Portland–South Portland–Biddeford, Maine metropolitan statistical area...

. In 1989, the refuge boundary expanded to include salt marsh
Salt marsh
A salt marsh is an environment in the upper coastal intertidal zone between land and salt water or brackish water, it is dominated by dense stands of halophytic plants such as herbs, grasses, or low shrubs. These plants are terrestrial in origin and are essential to the stability of the salt marsh...

, freshwater wetland
Wetland
A wetland is an area of land whose soil is saturated with water either permanently or seasonally. Wetlands are categorised by their characteristic vegetation, which is adapted to these unique soil conditions....

s and "critical edge" uplands around each of the nine divisions. In addition, the Biddeford Pool
Biddeford Pool
Biddeford Pool is the large tidal pool, located off Saco Bay south of the mouth of the Saco River on the south coast of Maine. It is approximately 6 miles southeast of downtown Biddeford, to which it is connected via State Route 208...

 Division, the tenth division of the refuge, was created. This division serves as a key staging area in southern Maine for a large number and diversity of shorebirds
Wader
Waders, called shorebirds in North America , are members of the order Charadriiformes, excluding the more marine web-footed seabird groups. The latter are the skuas , gulls , terns , skimmers , and auks...

. In 2007 the final Comprehensive Conservation Plan was signed, adding the eleventh division, York River
York River (Maine)
The York River is a stream in sout hiwest Maine, United States. It is tidal for over half of its length. It rises at York Pond in Eliot, and conjoined by brooks and creeks, feeds the tidal section...

 Division. When land acquisitions are complete, the refuge will be about 14,600 acres in size.

In 1984, a National Estuarine Research Reserve was established in Wells, Maine
Wells, Maine
Wells is a town in York County, Maine, United States. Founded in 1643, it is the third-oldest town in Maine. The population was 9,400 at the 2000 census. Wells Beach is a popular summer destination.-History:...

. The Reserve land is made up of portions of the Upper and Lower Wells divisions of the Refuge. Together, the Reserve and Refuge function to further the knowledge and understanding of estuaries throughout the community. The goal is to promote an increased stewardship
Stewardship
Stewardship is an ethic that embodies responsible planning and management of resources. The concept of stewardship has been applied in diverse realms, including with respect to environment, economics, health, property, information, and religion, and is linked to the concept of sustainability...

 and, ultimately, a greater protection of the estuaries.

Threatened and Endangered Wildlife Species

The Piping Plover is federally threatened and state endangered in Maine
Maine
Maine is a state in the New England region of the northeastern United States, bordered by the Atlantic Ocean to the east and south, New Hampshire to the west, and the Canadian provinces of Quebec to the northwest and New Brunswick to the northeast. Maine is both the northernmost and easternmost...

. Fifty to 75% of the Maine Piping Plover population nests at sites on or near the Refuge, including Crescent Surf Beach, Goosefare Brook, and Marshall Point at Goose Rocks
Goose Rocks
Goose Rocks Beach , is a coastal resort neighborhood located in the town of Kennebunkport, Maine USA, bordered by Cape Porpoise, Maine to the South, and Granite Point to the North...

.

New England Cottontail
New England Cottontail
The New England Cottontail is a species of cottontail rabbit represented by fragmented populations in areas of New England, specifically from southern Maine to southern New York...

 (Sylvilagus transitionalis) are found in Maine. Cottontails inhabit early successional
Ecological succession
Ecological succession, is the phenomenon or process by which a community progressively transforms itself until a stable community is formed. It is a fundamental concept in ecology, and refers to more or less predictable and orderly changes in the composition or structure of an ecological community...

 habitat that was relatively abundant in the early to mid-20th century. As farms were abandoned, the species did very well. Subsequently, increased development and reforestation
Reforestation
Reforestation is the natural or intentional restocking of existing forests and woodlands that have been depleted, usually through deforestation....

 has led to a population decline as this type of habitat became increasingly rare. The Refuge prohibited rabbiting
Rabbiting
Rabbiting is the sport of hunting rabbits, often using ferrets or dogs to track the prey and various trapping and shooting methods to catch them....

 starting in 1998 because of ongoing population declines. Subsequently, the Service was petitioned in 2000 to list the New England Cottontail under the Endangered Species Act
Endangered Species Act
The Endangered Species Act of 1973 is one of the dozens of United States environmental laws passed in the 1970s. Signed into law by President Richard Nixon on December 28, 1973, it was designed to protect critically imperiled species from extinction as a "consequence of economic growth and...

. Currently, New England Cottontail are listed as a Candidate Species under the Endangered Species Act and are endangered in Maine.

Salt marsh birds

In 1995, Sharp-tailed Sparrows were divided into two separate species: the Nelson's Sharp-tailed Sparrow and the Saltmarsh Sharp-tailed Sparrow. Saltmarsh Sharp-tailed Sparrows are found in salt marsh
Salt marsh
A salt marsh is an environment in the upper coastal intertidal zone between land and salt water or brackish water, it is dominated by dense stands of halophytic plants such as herbs, grasses, or low shrubs. These plants are terrestrial in origin and are essential to the stability of the salt marsh...

es along the Atlantic coast from the Delmarva Peninsula
Delmarva Peninsula
The Delmarva Peninsula is a large peninsula on the East Coast of the United States, occupied by most of Delaware and portions of Maryland and Virginia...

 north to southern Maine. Within the Refuge both species are found only on salt marshes. In fact, the Saltmarsh Sharp-tailed Sparrow is an obligate salt marsh species that spends its entire life cycle on salt marshes.

Waterbirds

Great Northern Loons (Gavia immer) frequent the lower reaches of tidal creeks of all Refuge Divisions from late autumn through early spring. They are commonly observed feeding on green crabs and small fish. During spring, summer and autumn migration, 11 species of wading birds
Wader
Waders, called shorebirds in North America , are members of the order Charadriiformes, excluding the more marine web-footed seabird groups. The latter are the skuas , gulls , terns , skimmers , and auks...

 use the estuarine systems of the Refuge.

Waterfowl

Twenty-six species of waterfowl are recorded from the Refuge. The most commonly observed species are American Black Duck
American Black Duck
The American Black Duck is a large dabbling duck. American Black Ducks are similar to Mallards in size, and resemble the female Mallard in coloration, although the Black Duck's plumage is darker...

 (Anas rubripes), Canada Goose
Canada Goose
The Canada Goose is a wild goose belonging to the genus Branta, which is native to arctic and temperate regions of North America, having a black head and neck, white patches on the face, and a brownish-gray body....

 (Branta canadensis), Mallard
Mallard
The Mallard , or Wild Duck , is a dabbling duck which breeds throughout the temperate and subtropical Americas, Europe, Asia, and North Africa, and has been introduced to New Zealand and Australia....

 (Anas platyrhynchos), Green-winged Teal
Green-winged Teal
The Green-winged Teal is a common and widespread duck that breeds in the northern areas of North America except on the Aleutian Islands. It was considered conspecific with the Common Teal The Green-winged Teal (Anas carolinensis) is a common and widespread duck that breeds in the northern areas of...

 (Anas carolinensis), Common Goldeneye
Common Goldeneye
The Common Goldeneye is a medium-sized sea duck of the genus Bucephala, the goldeneyes. Their closest relative is the similar Barrow's Goldeneye....

 (Bucephala clangula), Bufflehead
Bufflehead
The Bufflehead is a small American sea duck of the genus Bucephala, the goldeneyes. This species was first described by Linnaeus in his Systema naturae in 1758 as Anas albeola.-Description:...

 (Bucephala albeola), and Red-breasted Merganser
Red-breasted Merganser
The Red-breasted Merganser is a diving duck.-Taxonomy:The Red-breasted Merganser was one of the many species originally described by Linnaeus in his 18th-century work, Systema Naturae.-Description:...

 (Mergus serrator). Dabbling ducks
Anatinae
The Anatinae is a subfamily of the family Anatidae . Its surviving members are the dabbling ducks, which feed mainly at the surface rather than by diving...

 use salt pannes
Salt pannes and pools
Salt pannes and pools are water retaining depressions located within salt and brackish marshes. Pools tend to retain water during the summer months between high tides, whereas pannes generally do not. Salt pannes generally start when a mat of organic debris known as wrack, is deposited upon...

 and the upper reaches of tidal creeks, while diving duck
Diving duck
The diving ducks, commonly called pochards or scaups, are a category of duck which feed by diving beneath the surface of the water. They are part of the diverse and very large Anatidae family that includes ducks, geese, and swans....

s prefer deeper parts of the tidal creeks and the mouths of rivers and streams. Black Ducks, Mallards, and increasing numbers of Canada Geese breed on each Division of the Refuge. Wood Duck
Wood Duck
The Wood Duck or Carolina Duck is a species of duck found in North America. It is one of the most colourful of North American waterfowl.-Description:...

s (Aix sponsa) breed on the Upper Wells and Mousam River
Mousam River
The Mousam River is a river in Maine in the United States. Its source is Mousam Lake in York County, and it flows into the Atlantic Ocean just west of Kennebunk Beach. It flows through the towns of Sanford and Kennebunk.-External links:...

 Divisions each year.

Shorebirds

Southern coastal Maine is a migration and staging area for much of the North American shorebird population. Thousands of shorebirds feed along coastal beaches and mudflat
Mudflat
Mudflats or mud flats, also known as tidal flats, are coastal wetlands that form when mud is deposited by tides or rivers. They are found in sheltered areas such as bays, bayous, lagoons, and estuaries. Mudflats may be viewed geologically as exposed layers of bay mud, resulting from deposition of...

s as they migrate through the State. Biddeford Pool
Biddeford Pool
Biddeford Pool is the large tidal pool, located off Saco Bay south of the mouth of the Saco River on the south coast of Maine. It is approximately 6 miles southeast of downtown Biddeford, to which it is connected via State Route 208...

 serves as one of the top shorebird staging areas in southern Maine. The most common species observed in the autumn include Semipalmated Plover
Semipalmated Plover
The Semipalmated Plover is a small plover.This species weighs and measures in length and across the wings. Adults have a grey-brown back and wings, a white belly, and a white breast with one black neckband...

 (Charadrius semipalmatus), Black-bellied Plover
Grey Plover
The Grey Plover , known as the Black-bellied Plover in North America, is a medium-sized plover breeding in arctic regions. It is a long-distance migrant, with a nearly worldwide coastal distribution when not breeding....

 (Pluvialis squatarola), Least Sandpiper
Least Sandpiper
The Least Sandpiper is the smallest shorebird.This species has greenish legs and a short thin dark bill. Breeding adults are brown with dark brown streaks on top and white underneath. They have a light line above the eye and a dark crown. In winter, Least Sandpipers are grey above...

 (Calidris minutilla), Greater Yellowlegs
Greater Yellowlegs
The Greater Yellowlegs, Tringa melanoleuca, is a large North American shorebird, similar in appearance to the smaller Lesser Yellowlegs. Its closest relative, however, is the Greenshank, which together with the Spotted Redshank form a close-knit group...

 (Tringa melanoleuca), Short-billed Dowitcher
Short-billed Dowitcher
The Short-billed Dowitcher like its congener the Long-billed Dowitcher, is a medium-sized, stocky, long-billed shorebird in the family Scolopacidae. It is an inhabitant of North America, Middle America, and northern South America. It is strongly migratory; it completely vacates in breeding areas...

 (Limnodromus griseus), and Semipalmated Sandpiper
Semipalmated Sandpiper
The Semipalmated Sandpiper, Calidris pusilla, is a very small shorebird. It is sometimes separated with other "stints" in Erolia but although these apparently form a monophyletic group, the present species' old genus Ereunetes had been proposed before Erolia.Adults have black legs and a short stout...

 (Calidris pusilla). These species and others typically feed in the mudflats at low tide. Most shorebirds feed in salt pannes and roost in pannes and adjacent upland areas during high tides.

Gulls and terns

Herring Gulls (Larus argentatus) and Great Black-backed Gull
Great Black-backed Gull
The Great Black-backed Gull is the largest gull in the world, which breeds on the European and North American coasts and islands of the North Atlantic...

s (Larus marinus) are the most common gull species sighted on the refuge. They frequent all divisions throughout the year, but are most abundant in the autumn and winter when they roost on the marsh and tidal flats, and occasionally steal food from diving ducks in tidal creeks. Ring-billed Gull
Ring-billed Gull
The Ring-billed Gull is a medium-sized gull.Adults are length and with a wingspan. The head, neck and underparts are white; the relatively short bill is yellow with a dark ring; the back and wings are silver gray; and the legs are yellow. The eyes are yellow with red rims...

s (Larus delawarensis) also are common throughout the refuge, particularly during non-breeding season. During autumn and winter migration, Bonaparte's Gull
Bonaparte's Gull
The Bonaparte's Gull is a small gull.The Bonaparte's Gull is a small species, larger only than the Little Gull and the Saunders's Gull among all gull species. Adults are long with a wingspan and a body mass of . They have a black hood and a short thin dark bill. The body is mainly white with...

s (Larus philadelphia) feed and roost at the mouths of tidal creeks and rivers throughout the refuge, but they are most abundant on the Biddeford Pool, Upper Wells, and Lower Wells divisions.

Least Terns nest on the refuge in several locations. In the mid-1980s, Common Tern
Common Tern
The Common Tern is a seabird of the tern family Sternidae. This bird has a circumpolar distribution, breeding in temperate and sub-Arctic regions of Europe, Asia and east and central North America. It is strongly migratory, wintering in coastal tropical and subtropical regions. It is sometimes...

s nested in the salt marsh on the Lower Wells and Little River divisions. Roseate Tern
Roseate Tern
The Roseate Tern is a seabird of the tern family Sternidae. This bird has a number of geographical races, differing mainly in bill colour and minor plumage details....

s (Sterna dougallii) nested on West Goose Rocks Island in 1985, and lately, have been observed along Crescent Surf Beach in the Upper Wells Division. In 2003, Crescent Surf Beach hosted the largest nesting colony (157 pairs) of Least Terns in Maine. Early season crow predation and late season owl and coyote predation depressed productivity. The Refuge controls diurnal predators such as crows and foxes with several techniques, including hazing, fencing, trapping, and shooting. Least Terns also nest at Laudholm Beach, Goose Rocks
Goose Rocks
Goose Rocks Beach , is a coastal resort neighborhood located in the town of Kennebunkport, Maine USA, bordered by Cape Porpoise, Maine to the South, and Granite Point to the North...

, Higgins
Higgins Beach
Higgins Beach is a small beach located in the state of Maine in the United States. It is located in the town of Scarborough in Cumberland County. The beach is north of Prouts Neck and Old Orchard Beach and south of Crescent Beach State Park....

, and Reid State Park
Reid State Park
Reid State Park is a state park on Georgetown Island, in Sagadahoc County, Maine. There are several naturally sandy beaches which contrast with the usually rocky coast of Maine. The conditions make for excellent swimming during the summer months, with a natural inlet providing a somewhat warmer...

. During migration, large numbers of Common Terns, along with smaller numbers of Roseate Terns (15), stage at Crescent Surf Beach

Land birds

Over 120 species of land birds have been recorded at the refuge with over 72 nesting. Some of those include Great Crested Flycatcher
Great Crested Flycatcher
The Great Crested Flycatcher is a large insect-eating bird of the tyrant flycatcher family. It is the most widespread member of the genus, Myiarchus, in North America and is found over most of the eastern and mid-western portions of the continent...

 (Myiarchus crinitus), Tree Swallow
Tree Swallow
The Tree Swallow, Tachycineta bicolor, is a migratory passerine bird that breeds in North America and winters in Mexico, Central America and the Caribbean. It is a very rare vagrant to western Europe....

 (Tachycineta bicolor), Hermit Thrush
Hermit Thrush
The Hermit Thrush is a medium-sized North American thrush. It is not very closely related to the other North American migrant species of Catharus, but rather to the Mexican Russet Nightingale-thrush.-Description:...

 (Catharus guttatus), Black-throated Green Warbler
Black-throated Green Warbler
The Black-throated Green Warbler, Setophaga virens, is a small songbird of the New World warbler family.It is 12 cm long and weighs 9 g, and has an olive-green crown, a yellow face with olive markings, a thin pointed bill, white wing bars, an olive-green back and pale underparts with...

 (Dendroica virens), Scarlet Tanager
Scarlet Tanager
The Scarlet Tanager is a medium-sized American songbird. Formerly placed in the tanager family , it and other members of its genus are now classified in the cardinal family . The species's plumage and vocalizations are similar to other members of the cardinal family.-Description:Adults have pale...

 (Piranga olivacea), and others. Forests, woodlands, and swamps surrounding Refuge salt marshes also provide habitat for many raptors
Bird of prey
Birds of prey are birds that hunt for food primarily on the wing, using their keen senses, especially vision. They are defined as birds that primarily hunt vertebrates, including other birds. Their talons and beaks tend to be relatively large, powerful and adapted for tearing and/or piercing flesh....

. Many migrating raptors use forested areas next to marshes as hunting perches and feeding areas. Sharp-shinned Hawk
Sharp-shinned Hawk
The Sharp-shinned Hawk is a small hawk. In fact, "sharp-shins" or "sharpies" are the smallest to reside in USA and Canada, though some Neotropical species are smaller...

 (Accipiter striatus), Cooper's Hawk
Cooper's Hawk
Cooper's Hawk is a medium-sized hawk native to the North American continent and found from Canada to Mexico. As in many birds of prey, the male is smaller than the female...

 (Accipiter cooperii), and Broad-winged Hawk
Broad-winged Hawk
The Broad-winged Hawk is a small hawk of the genus Buteo. During the summer some subspecies are distributed over eastern North America, as far west as British Columbia and Texas; they then migrate south to winter in the neotropics from Mexico down to southern Brazil. Other subspecies are all-year...

 (Buteo platypterus) have nested in forested habitat on the Refuge. Northern Goshawks (Accipiter gentilis) and Red-tailed Hawk
Red-tailed Hawk
The Red-tailed Hawk is a bird of prey, one of three species colloquially known in the United States as the "chickenhawk," though it rarely preys on standard sized chickens. It breeds throughout most of North America, from western Alaska and northern Canada to as far south as Panama and the West...

s (Buteo jamaicensis) nest in the area. During migration (primarily autumn), many raptors move through the Refuge. Northern Harriers
Hen Harrier
The Hen Harrier or Northern Harrier is a bird of prey. It breeds throughout the northern parts of the northern hemisphere in Canada and the northernmost USA, and in northern Eurasia. This species is polytypic, with two subspecies. Marsh Hawk is a historical name for the American form.It migrates...

 are the only raptor species thought to breed in the estuarine
Estuary
An estuary is a partly enclosed coastal body of water with one or more rivers or streams flowing into it, and with a free connection to the open sea....

 communities of the Refuge.

Mammals

White-tailed deer
White-tailed Deer
The white-tailed deer , also known as the Virginia deer or simply as the whitetail, is a medium-sized deer native to the United States , Canada, Mexico, Central America, and South America as far south as Peru...

 (Odocoileus virginianus) are the mammal
Mammal
Mammals are members of a class of air-breathing vertebrate animals characterised by the possession of endothermy, hair, three middle ear bones, and mammary glands functional in mothers with young...

 most commonly observed on the refuge. Their trails cut through certain portions of the salt marsh
Salt marsh
A salt marsh is an environment in the upper coastal intertidal zone between land and salt water or brackish water, it is dominated by dense stands of halophytic plants such as herbs, grasses, or low shrubs. These plants are terrestrial in origin and are essential to the stability of the salt marsh...

 on each division, although they more typically are observed along marsh edges and in surrounding forests. The refuge lies entirely within Wildlife Management District No. 24, which had an estimated winter population of 30 deer/mi2 in 1997. The Wildlife Division Research and Management Report (2000) stated that the herd has continued to grow at 15 percent per year, and the wintering population is now nearly 40 deer/mi2. In certain areas of the refuge, hunting (including white-tailed deer) is prohibited because of state designated Game Sanctuaries. Deer populations in those areas are estimated from 50 to more than 100 deer/mi2. Those populations far exceed the state target of 50 percent to 60 percent of carrying capacity
Carrying capacity
The carrying capacity of a biological species in an environment is the maximum population size of the species that the environment can sustain indefinitely, given the food, habitat, water and other necessities available in the environment...

.

Raccoon
Raccoon
Procyon is a genus of nocturnal mammals, comprising three species commonly known as raccoons, in the family Procyonidae. The most familiar species, the common raccoon , is often known simply as "the" raccoon, as the two other raccoon species in the genus are native only to the tropics and are...

 (Procyon lotor) tracks and scats abound on all divisions. Their sign most often appears along the edges of tidal creeks and salt pannes, where they search for green crabs and small fish. Care must be taken to distinguish raccoon sign from that of river otters (Lutra canadensis), another mammal that forages extensively in the marshes, and is infrequently observed in the salt hay
Spartina patens
Spartina patens , also known as salt marsh hay, is a species of cordgrass native to the Atlantic coast of the Americas, from Newfoundland south along the eastern United States to the Caribbean and northeast Mexico...

 along the edges of tidal creeks. Most recently, river otters were seen in the Merriland River
Merriland River
The Merriland River is a river in southern Maine. It is a tributary of the Little River, a tidal river flowing to the Atlantic Ocean.The Merriland River rises in the southern corner of Sanford and flows east into Wells, which it crosses to the east border of the town and ends at the Little...

 and Branch Brook in the Upper Wells Division.

Mink
American Mink
The American mink is a semi-aquatic species of Mustelid native to North America, though human intervention has expanded its range to many parts of Europe and South America. Because of this, it is classed as Least Concern by the IUCN. Since the extinction of the sea mink, the American mink is the...

 (Mustela vison), Striped Skunk
Striped Skunk
The striped skunk, Mephitis mephitis, is an omnivorous mammal of the skunk family Mephitidae. Found over most of the North American continent north of Mexico, it is one of the best-known mammals in Canada and the United States.-Description:...

 (Mephitis mephitis), Red Fox
Red Fox
The red fox is the largest of the true foxes, as well as being the most geographically spread member of the Carnivora, being distributed across the entire northern hemisphere from the Arctic Circle to North Africa, Central America, and the steppes of Asia...

 (Vulpes fulva), and Coyote
Coyote
The coyote , also known as the American jackal or the prairie wolf, is a species of canine found throughout North and Central America, ranging from Panama in the south, north through Mexico, the United States and Canada...

 (Canis latrans) also hunt in the estuary. Beaver (Castor canadensis) and Muskrat
Muskrat
The muskrat , the only species in genus Ondatra, is a medium-sized semi-aquatic rodent native to North America, and introduced in parts of Europe, Asia, and South America. The muskrat is found in wetlands and is a very successful animal over a wide range of climates and habitats...

 (Ondatra zibethicus) are occasionally seen swimming in tidal creeks. A few Harbor seal
Harbor Seal
The harbor seal , also known as the common seal, is a true seal found along temperate and Arctic marine coastlines of the Northern Hemisphere...

 (Phoca vitulina) haul-out sites exist on the Brave Boat Harbor, Lower Wells, Mousam River, and Goose Rocks divisions. Peak use occurs during the winter, but individuals are observed throughout the year. The Lower Wells haul-out site receives the most use, with peak counts of 30 seals. During the winter months Harp Seal
Harp Seal
The harp seal or saddleback seal is a species of earless seal native to the northernmost Atlantic Ocean and adjacent parts of the Arctic Ocean. It now belongs to the monotypic genus Pagophilus. Its scientific name, Pagophilus groenlandicus, means "ice-lover from Greenland", and its synonym, Phoca...

s (Pagophilus groenlandicus), and occasionally Hooded
Hooded Seal
The hooded seal is an arctic pinniped found only in the central and western North Atlantic ranging from Svalbard in the east to the Gulf of St...

 (Cystophora cristata) and Grey Seal
Grey Seal
The grey seal is found on both shores of the North Atlantic Ocean. It is a large seal of the family Phocidae or "true seals". It is the only species classified in the genus Halichoerus...

s (Halichoerus grypus), can be found basking on refuge salt marshes and in offshore waters. Seal strandings are a common occurrence, and are reported to marine animal rescue agencies.

Many large mammals are found on or near the refuge. Moose
Moose
The moose or Eurasian elk is the largest extant species in the deer family. Moose are distinguished by the palmate antlers of the males; other members of the family have antlers with a dendritic configuration...

 (Alces alces) and Black Bear
American black bear
The American black bear is a medium-sized bear native to North America. It is the continent's smallest and most common bear species. Black bears are omnivores, with their diets varying greatly depending on season and location. They typically live in largely forested areas, but do leave forests in...

 (Ursus americanus) are becoming more common in southern Maine as their populations continue to grow. They have been sighted on all Refuge divisions except Moody. A bobcat
Bobcat
The bobcat is a North American mammal of the cat family Felidae, appearing during the Irvingtonian stage of around 1.8 million years ago . With twelve recognized subspecies, it ranges from southern Canada to northern Mexico, including most of the continental United States...

 (Lynx rufus) was reported as sporadically using the Upper and Lower Wells divisions in 1991 and 1992. Fishers
Fisher (animal)
The fisher is a medium-size mammal native to North America. It is a member of the mustelid family, commonly referred to as the weasel family. The fisher is closely related to but larger than the American Marten...

 (Martes pennanti) are increasingly sighted on the refuge; a vehicle killed a fisher near refuge headquarters in 1998, and several sightings around the headquarters have occurred since then. Gray fox
Gray Fox
The gray fox is a mammal of the order Carnivora ranging throughout most of the southern half of North America from southern Canada to the northern part of South America...

 (Urocyon cinereoargenteus) and short-tailed weasel
Stoat
The stoat , also known as the ermine or short-tailed weasel, is a species of Mustelid native to Eurasia and North America, distinguished from the least weasel by its larger size and longer tail with a prominent black tip...

 (Mustela erminea) most likely use several refuge divisions. Porcupines
North American Porcupine
The North American Porcupine , also known as Canadian Porcupine or Common Porcupine, is a large rodent in the New World porcupine family. The Beaver is the only rodent larger than the North American Porcupine found in North America...

 (Erethizon dorsatum) and woodchucks
Groundhog
The groundhog , also known as a woodchuck, whistle-pig, or in some areas as a land-beaver, is a rodent of the family Sciuridae, belonging to the group of large ground squirrels known as marmots. Other marmots, such as the yellow-bellied and hoary marmots, live in rocky and mountainous areas, but...

 (Marmota monax) are found throughout the refuge, where they occur in varied habitats.

Snowshoe Hare
Snowshoe Hare
The Snowshoe Hare , also called the Varying Hare, or Snowshoe Rabbit, is a species of hare found in North America. It has the name "snowshoe" because of the large size of its hind feet and the marks its tail leaves. The animal's feet prevent it from sinking into the snow when it hops and walks...

 (Lepus americanus) are found in forests throughout the refuge in areas with dense understory. White-footed Mice
White-footed mouse
White-footed Mouse is a rodent native to North America. It ranges from Ontario, Quebec, Labrador and the Maritime Provinces to the southwest USA and Mexico. It is also known as the Woodmouse, particularly in Texas.Adults are in length, not counting the tail, which can add another . A young adult...

 (Peromyscus leucopus), meadow jumping mice
Meadow jumping mouse
The meadow jumping mouse is the most widely distributed mouse in the subfamily Zapodinae. It may be found from the Atlantic coast, to the Great Plains, as far north as the arctic tree lines in Canada and Alaska, and as far south as Georgia, Alabama, Arizona, and New Mexico.- Description :The...

 (Zapus hudsonius), and Meadow Vole
Meadow Vole
The Meadow Vole , sometimes called the Field Mouse or Meadow Mouse, is a North American vole found across Canada, Alaska and the northern United States. Its range extends further south along the Atlantic coast. One subspecies, the Florida Salt Marsh Vole , is found in Florida, and is classified as...

s (Microtus pennsylvanicus) occasionally use the edge of salt marsh habitat. Masked Shrews (Sorex cinereus), Northern Short-tailed Shrew
Northern Short-tailed Shrew
The Northern Short-tailed Shrew is the largest shrew in the genus Blarina, and occurs in the northeastern region of North America. It is a semifossorial, highly active and voracious insectivore and is present in a variety of habitats. It is notable in that it is one of the few venomous mammals...

s (Blarina brevicauda), Southern Red-backed Vole
Southern Red-backed Vole
The Southern Red-backed Vole or Gapper's Red-backed Vole is a small slender vole found in Canada and the northern United States...

s (Clethrionomys gapperi), and Pine Voles
Woodland Vole
The Woodland Vole, Microtus pinetorum, is a small vole found in eastern North America. It is also known as the Pine Vole.-Description:...

 (Microtus pinetorum) have also been caught in salt marshes.

Other small mammals that commonly are found on the refuge include eastern chipmunk
Eastern Chipmunk
The eastern chipmunk is a small squirrel-like rodent found in eastern North America, the sole living member of the chipmunk genus and subgenus Tamias....

 (Tamias striatus), American Red Squirrel
American Red Squirrel
The American Red Squirrel is one of three species of tree squirrel currently classified in the genus Tamiasciurus and known as pine squirrels...

 (Tamiasciurus hudsonicus), and eastern grey squirrel (Sciurus carolinensis). These species are most common in pine-oak forests where acorns are abundant. Southern flying squirrel
Southern Flying Squirrel
The Southern Flying Squirrel is one of two species of the genus Glaucomys, the only flying squirrels found in North America . It is found in deciduous and mixed woods in the eastern half of North America, from southeastern Canada, to Florida, USA...

 (Glaucomys volans) is recorded for the Upper Wells and Brave Boat divisions, but they probably also occur in other areas with mature pine-oak forest. Other small mammals that are known or are likely to occur on the refuge include Hairy-tailed Mole
Hairy-tailed Mole
The Hairy-tailed Mole , also known as Brewer's Mole, is a medium-sized North American mole. It is the only member of the genus Parascalops....

 (Parascalops breweri), star-nosed mole
Star-nosed mole
The star-nosed mole is a small mole found in wet low areas of eastern Canada and the north-eastern United States, with records extending along the Atlantic coast as far as extreme southeastern Georgia...

 (Condylura cristata), Smoky Shrew
Smoky Shrew
The Smoky Shrew, Sorex fumeus, is a medium-sized North American shrew found in eastern Canada and the northeastern United States.It is dull grey in colour with lighter underparts and a long tail which is brown on top and yellowish underneath. During winter, its fur is grey...

 (Sorex fumeus), and house mouse
House mouse
The house mouse is a small rodent, a mouse, one of the most numerous species of the genus Mus.As a wild animal the house mouse mainly lives associated with humans, causing damage to crops and stored food....

 (Mus musculus).

Reptiles and amphibians

The refuge has a limited amount of freshwater cattail
Typha
Typha is a genus of about eleven species of monocotyledonous flowering plants in the family Typhaceae. The genus has a largely Northern Hemisphere distribution, but is essentially cosmopolitan, being found in a variety of wetland habitats...

 marsh or pond habitat. However, within its
uplands, the refuge protects an extensive network of rivers, uplands and vernal pool
Vernal pool
Vernal pools, also called vernal ponds or ephemeral pools, are temporary pools of water. They are usually devoid of fish, and thus allow the safe development of natal amphibian and insect species...

s, which provide important amphibian
Amphibian
Amphibians , are a class of vertebrate animals including animals such as toads, frogs, caecilians, and salamanders. They are characterized as non-amniote ectothermic tetrapods...

 and reptile
Reptile
Reptiles are members of a class of air-breathing, ectothermic vertebrates which are characterized by laying shelled eggs , and having skin covered in scales and/or scutes. They are tetrapods, either having four limbs or being descended from four-limbed ancestors...

 habitat. Anuran call counts and limited vernal pool surveys were conducted on the refuge. American toad
American toad
The American Toad is a common species of toad found throughout the eastern United States and Canada. It is divided into three subspecies—the Eastern American Toad , the Dwarf American Toad , and the rare Hudson Bay Toad...

 (Bufo americanus), green frog (Rana clamitans), Wood Frog
Wood Frog
The Wood Frog has a broad distribution over North America, extending from the southern Appalachians to the boreal forest with several notable disjunct populations including lowland eastern North Carolina...

 (Rana sylvatica), Pickerel Frog
Pickerel Frog
The Pickerel Frog is a small North American frog, characterized by the appearance of seemingly "hand-drawn" squares on its dorsal surface.-Distinguishing features:...

 (Rana palustris), Bullfrog
Bullfrog
The American bullfrog , often simply known as the bullfrog in Canada and the United States, is an aquatic frog, a member of the family Ranidae, or “true frogs”, native to much of North America. This is a frog of larger, permanent water bodies, swamps, ponds, and lakes, where it is usually found...

 (Rana catesbeiana), gray tree frog (Hyla versicolor) and Spring Peeper
Spring Peeper
The Spring Peeper is a small chorus frog widespread throughout the eastern USA and Canada.-Subspecies:There are two subspecies of the Spring Peeper, the Northern and the Southern Spring Peeper . The Northern is similar to the Southern except for a strong dark marking on the Southern frog's belly...

 (Pseudacris crucifer) are documented as breeding on most refuge divisions. In addition, yellow-spotted salamanders
Spotted Salamander
The Spotted Salamander or Yellow-spotted Salamander is a mole salamander common in the eastern United States and Canada. The Spotted Salamander is the State amphibian of South Carolina. It has recently been found that its embryos have algae living inside them in a mutualistic...

 (Ambystoma maculatum), Red Back Salamander
Red Back Salamander
The red back salamander is a small, hardy woodland salamander. It inhabits wooded slopes in Eastern North America; west to Missouri; south to North Carolina; and north from southern Quebec and the Maritime Provinces in Canada to Minnesota...

s (Plethodon cinereus) and eastern newt
Eastern Newt
The Eastern Newt or Red-spotted Newt is a common salamander of eastern North America. Eastern newts dwell in wet forests with small lakes or ponds. They can coexist in an aquatic environment with fish, however, their skin secretes a poisonous substance when the newt is threatened or injured...

s (Notophthalmus viridescens) are recorded as common breeders. The Blue-spotted Salamander
Blue-spotted Salamander
The Blue-spotted salamander, or Ambystoma laterale, is a mole salamander native to the Great Lakes states and northeastern United States, and parts of Ontario and Quebec in Canada. Their range is known to extend to James Bay to the north, and southeastern Manitoba to the west.-Description:These...

 (Ambystoma laterale) and Northern Leopard Frog
Northern Leopard Frog
The Northern Leopard Frog is a species of leopard frog from the true frog family, native to parts of Canada and United States. It is the state amphibian of Minnesota and Vermont.-Physical description:...

 (Rana pipiens) are uncommon, but likely are breeders on the refuge.

Documented species on the Refuge

Garter snake
Common Garter Snake
The Common Garter Snake is a snake indigenous to North America. Most garter snakes have a pattern of yellow stripes on a brown or green background and their average length is about , maximum about .-Subspecies:...

 (Thamnophis sirtalis), ribbon snake
Ribbon snake
The Ribbon Snake or Ribbonsnake is a common snake found throughout North America. It averages in length and is a member of the garter snake genus...

 (Thamnophis sauritus, Maine—Special Concern), smooth green snake (Liochlorophis vernalis), redbelly snake
Storeria occipitomaculata
The redbelly snake is a species of snake found in North America. The following subspecies are known:*Florida redbelly snake - Storeria occipitomaculata obscura Trapido, 1944...

 (Storeria occipitomaculata), painted turtle
Painted Turtle
The painted turtle is the most widespread native turtle of North America. It lives in slow-moving fresh waters, from southern Canada to Louisiana and northern Mexico, and from the Atlantic to the Pacific. The turtle is the only species of the genus Chrysemys, which is part of the pond turtle...

 (Chrysemys picta), snapping turtle (Chelydra serpentina) and spotted turtle
Spotted Turtle
The Spotted turtle , the only current species of Clemmys, is a small, semi-aquatic turtle that reaches a carapace length of upon adulthood. Their broad, smooth, low dark-colored upper shell, or carapace, ranges in its exact colour from black to a bluish black with a number of yellow tiny round spots...

 (Clemmys guttata, Maine—Threatened). Species that are likely to use the refuge but are not documented include ringneck snake
Diadophis punctatus
The ringneck snake or ring-necked snake is a colubrid snake species. It is found throughout much of the United States, central Mexico, and south eastern Canada. Ring-necked snakes are secretive, nocturnal snakes that are rarely seen during the day time...

 (Diadophis punctatus), Milk Snake
Milk Snake
The milk snake or milksnake is a species of king snake. There are 25 subspecies among the milk snakes, including the commonly named scarlet kingsnake...

 (Lampropeltis triangulum), Northern Water Snake
Northern Water Snake
The Northern water snake is a large, nonvenomous, well-known snake in the Colubridae family that is native to North America.-Behavior:...

 (Nerodia sipedon), brown snake
Storeria dekayi
Storeria dekayi, commonly known as the brown snake or De Kay's snake, is a small species of colubrid snake.-Geographic range:...

 (Storeria dekayi, Maine Special Concern), Blanding's turtle
Blanding's Turtle
Blanding's turtle is a semi-aquatic turtle of the family Emydidae. It is considered to be an endangered species throughout much of its range.-Taxonomy:...

 (Emydoidea blandingii, Maine Endangered) and possibly, eastern racer (Coluber constrictor
Coluber constrictor
Coluber constrictor is a species of nonvenomous, colubrid snakes commonly referred to as the eastern racers. They are primarily found throughout the United States, east of the Rocky Mountains, but they range north into Canada, and south into Mexico, Guatemala and Belize...

, Maine— Endangered), wood turtle
Wood Turtle
The wood turtle is a turtle endemic to North America. It is in the genus Glyptemys, a designation given to only one other turtle: the bog turtle. The wood turtle reaches a carapace length of , its defining characteristic being the pyramidal pattern on its upper shell...

 (Clemmys insculpta, Maine—Special Concern) and common musk turtle
Sternotherus odoratus
Sternotherus odoratus is a species of small turtle native to southeastern Canada and much of the Eastern United States. It is also known as the common musk turtle or stinkpot due to its ability to release a foul musky odor to deter predation...

 (Sternotherus odoratus). Records indicate that both Blanding’s turtle and spotted turtle occur in many locations along the refuge boundary. Wood turtle and black racer records are much less common, and musk turtle records in the vicinity of the refuge are nonexistent. Surveys targeted at detecting turtles and snakes should be developed and implemented on refuge lands with particular attention to the occurrence of the rare, secretive Blanding’s turtle. Lands within the proposed acquisition boundary in Kennebunk and Biddeford have extensive vernal pool habitat that will benefit several Species of Concern
Species of Concern
In wildlife conservation, Species of Concern is an informal term, not defined in the federal Endangered Species Act. The term commonly refers to species that are declining or appear to be in need of concentrated conservation actions. Many agencies and organizations maintain lists of these at-risk...

.

Fish

Coastal marsh
Marsh
In geography, a marsh, or morass, is a type of wetland that is subject to frequent or continuous flood. Typically the water is shallow and features grasses, rushes, reeds, typhas, sedges, other herbaceous plants, and moss....

es, bays, tidal creeks
Creek (tidal)
A tidal creek, tidal channel, or estuary is the portion of a stream that is affected by ebb and flow of ocean tides, in the case that the subject stream discharges to an ocean, sea or strait. Thus this portion of the stream has variable salinity and electrical conductivity over the tidal cycle...

 and rivers support diverse shellfish and finfish populations. Sunfish (Lepomis
Lepomis
Lepomis is a genus of freshwater fish in the sunfish family of order Perciformes. The most recognizable species of the sixteen in this genus is the Bluegill....

 spp.
), creek chub
Semotilus atromaculatus
Semotilus atromaculatus is a small minnow found in the eastern two-thirds of the US and eastern Canada...

 (Semotilus atromaculatus), cunner
Bergall
A bergall or cunner or conner, Tautogolabrus adspersus, is a saltwater fish found in the western Atlantic. It is edible and its musky taste is considered a delicacy by some. Specimens may grow to 12 inches though most range from 4-10. Often bergalls are found mixed in with black fish , living...

 (Tautogolabrus adspersus), golden shiner
Golden shiner
The golden shiner is a cyprinid fish native to eastern North America. It is the sole member of its genus. Much used as a bait fish, it is probably the most widely pond-cultured fish in the United States.-Description:...

 (Notemigonus crysoleucas), common mummichog
Mummichog
The mummichog is a killifish also known as mummies, gudgeons, and mud minnows are found in brackish and coastal waters along the eastern seaboard of the United States as well as Atlantic Canada. It is noted for its hardiness and for being a popular research subject in embryological, physiological,...

, American eel
American eel
The American eel, Anguilla rostrata, is a catadromous fish found on the eastern coast of North America. It has a snake-like body with a small sharp pointed head. It is brown on top and a tan-yellow color on the bottom. It has sharp pointed teeth but no pelvic fins...

 (Anguilla rostrata) and White Sucker
White Sucker
The White Sucker is a bottom-feeding freshwater fish inhabiting North America from Labrador in the north to Georgia and New Mexico in the south. It is a long, round-bodied fish with a dark green, grey, copper, brown, or black back and sides and a light underbelly. When fullgrown, it is between 12...

 (Catostomus commersoni) abound. Brook trout
Brook trout
The brook trout, Salvelinus fontinalis, is a species of fish in the salmon family of order Salmoniformes. In many parts of its range, it is known as the speckled trout or squaretail. A potamodromous population in Lake Superior are known as coaster trout or, simply, as coasters...

 (Salvelinus fontinalis) and brown trout
Brown trout
The brown trout and the sea trout are fish of the same species....

 (Salmo trutta) are stocked in rivers and estuaries each year. The Ogunquit River
Ogunquit River
The Ogunquit River is a river in southern Maine. It flows to the Atlantic Ocean at the town of Ogunquit.-References:**...

 sustains alewife
Alewife
The alewife is a species of herring. There are anadromous and landlocked forms. The landlocked form is also called a sawbelly or mooneye...

 (Alosa pseudoharengus), blueback herring
Blueback Shad
Blueback Shad is a species of ray-finned fish in the genus Alosa. These fish are silvery in color, have a series of scutes along their belly, and are characterized by deep bluish green backs...

 (Alosa aestivalis), pollock
Pollachius virens
Pollachius virens is a species of marine fish in the Pollachius genus. Together with Pollachius pollachius it is generally referred to in the U.S. as Pollock. Other names include the Boston blues , coalfish and saithe in the UK.This species can be separated from P...

 (Pollachius virens), bluefish
Bluefish
The bluefish , called tailor in Australia, is a species of popular marine gamefish found in all climates. It is the sole species of the Pomatomidae family....

 (Pomatomus saltatrix), longhorn sculpin (Myoxocephalus octodecimspinosus), and winter flounder
Winter flounder
The winter flounder, Pseudopleuronectes americanus, is a right-eyed flatfish of the family Pleuronectidae. It is native to coastal waters of the western north Atlantic coast, from Labrador, Canada to Georgia, United States...

 (Pleuronectes americanus). The Webhannet River
Webhannet River
The Webhannet River is an river whose watershed is contained entirely within the town of Wells, Maine.The river has five tributaries, including three with official names: Pope’s Creek, Depot Brook, and Blacksmith Brook...

 has native species such as winter flounder, northern pipefish
Northern pipefish
Northern pipefish is a species of the pipefishes, which widespread in the north-western Atlantic from the Gulf of Saint Lawrence in Canada to north-eastern Florida in USA and north-western Gulf of Mexico. Marine \ freshwater demersal amphidromous fish, up to 30.0 cm length. Found on the depth...

 (Syngnathus fuscus), Atlantic herring
Atlantic herring
Atlantic herring is a fish in the family Clupeidae. It is one of the most abundant fish species on earth. Herring can be found on both sides of the Atlantic Ocean, congregating in large schools. They can grow up to in length and weigh more than...

 (Clupea harengus), common mummichog, Atlantic silverside
Atlantic silverside
The Atlantic silverside, Menidia menidia, is one of the most common fish in the Chesapeake Bay and in the Barnegat Bay. It is a common subject of scientific research because has a sensitivity to environmental changes....

s (Menidia menidia) and Atlantic mackerel
Atlantic mackerel
The Atlantic mackerel , is a pelagic schooling species of mackerel found on both sides of the North Atlantic Ocean. The species is also called Boston mackerel, or just mackerel....

. The Merriland River
Merriland River
The Merriland River is a river in southern Maine. It is a tributary of the Little River, a tidal river flowing to the Atlantic Ocean.The Merriland River rises in the southern corner of Sanford and flows east into Wells, which it crosses to the east border of the town and ends at the Little...

 sustains populations of American eel, brown trout, and brook trout. The Mousam River attracts little skate
Little skate
The little skate, Raja erinacea, is a species of skate in the family Rajidae, found from Nova Scotia to North Carolina on sand or gravel habitats. They are one of the dominant members of the demersal fish community in the northwestern Atlantic...

 (Raja erinacea), American shad
American shad
-Introduction:The American shad or Atlantic shad, Alosa sapidissima, is a species of anadromous fish in family Clupeidae of order Clupeiformes. It is not closely related to the other North American shads...

 (Alosa sapidissima), striped bass
Striped bass
The striped bass is the state fish of Maryland, Rhode Island, South Carolina, and the state saltwater fish of New York, Virginia, and New Hampshire...

 (Morone saxatilis), bluefish, cunner, Atlantic mackerel (Scomber scombrus), pollock, and rainbow smelt
Rainbow smelt
The rainbow smelt, Osmerus mordax, is an anadromous species of fish of the family Osmeridae. The distribution of Osmerus mordax is circumpolar. The rainbow smelt was introduced to the Great Lakes, and from there has made its way to various other places. Walleye, trout, and other larger fish prey on...

 (Osmerus mordax). The Spurwink River
Spurwink River
The Spurwink River is a primarily tidal river in Cumberland County, Maine, in the United States. It rises in the town of Cape Elizabeth and flows west, then southwest, through salt marshes to its mouth at the Atlantic Ocean. It is bridged by Maine State Route 77 at the river's approximate halfway...

 supports blueback herring, Atlantic menhaden
Atlantic menhaden
The Atlantic menhaden is a silvery, highly compressed fish in the herring family, Clupeidae. A filter feeder, it lives on plankton caught in midwater. Adult fish can filter up to four gallons of water a minute; and they play an important role in clarifying ocean water...

 (Brevoortia tyrannus), American shad, pollock, cunner, winter flounder, and little skate. Striped bass and brown trout are popular recreational fishing resources in the area. The National Marine Fisheries Service
National Marine Fisheries Service
The National Marine Fisheries Service is a United States federal agency. A division of the National Oceanic and Atmospheric Administration and the Department of Commerce, NMFS is responsible for the stewardship and management of the nation's living marine resources and their habitat within the...

 has designated as “essential fish habitat” areas that provide substrate necessary for fish spawning
Spawn (biology)
Spawn refers to the eggs and sperm released or deposited, usually into water, by aquatic animals. As a verb, spawn refers to the process of releasing the eggs and sperm, also called spawning...

, breeding, feeding, or growth to maturity. Estuaries within the refuge boundaries are part of that essential fish habitat.

Habitat types

Rachel Carson NWR is approximately 35 percent tidal, 10 percent freshwater wetlands and 55 percent uplands. Tidal habitat
Habitat
* Habitat , a place where a species lives and grows*Human habitat, a place where humans live, work or play** Space habitat, a space station intended as a permanent settlement...

s include beach, dune
Dune
In physical geography, a dune is a hill of sand built by wind. Dunes occur in different forms and sizes, formed by interaction with the wind. Most kinds of dunes are longer on the windward side where the sand is pushed up the dune and have a shorter "slip face" in the lee of the wind...

, dune grassland, river, rocky shore, estuarine, bay and salt marsh
Salt marsh
A salt marsh is an environment in the upper coastal intertidal zone between land and salt water or brackish water, it is dominated by dense stands of halophytic plants such as herbs, grasses, or low shrubs. These plants are terrestrial in origin and are essential to the stability of the salt marsh...

. Freshwater wetlands include cattail marsh, bog
Bog
A bog, quagmire or mire is a wetland that accumulates acidic peat, a deposit of dead plant material—often mosses or, in Arctic climates, lichens....

, emergent scrub-shrub wetlands, pocket swamps, red maple swamps and floodplain
Floodplain
A floodplain, or flood plain, is a flat or nearly flat land adjacent a stream or river that stretches from the banks of its channel to the base of the enclosing valley walls and experiences flooding during periods of high discharge...

 forest. The majority of the upland forests consist of mixed oak
Oak
An oak is a tree or shrub in the genus Quercus , of which about 600 species exist. "Oak" may also appear in the names of species in related genera, notably Lithocarpus...

 and pine
Pine
Pines are trees in the genus Pinus ,in the family Pinaceae. They make up the monotypic subfamily Pinoideae. There are about 115 species of pine, although different authorities accept between 105 and 125 species.-Etymology:...

 forest, however hemlock
Tsuga
Tsuga is a genus of conifers in the family Pinaceae. The common name hemlock is derived from a perceived similarity in the smell of its crushed foliage to that of the unrelated plant poison hemlock....

, spruce
Spruce
A spruce is a tree of the genus Picea , a genus of about 35 species of coniferous evergreen trees in the Family Pinaceae, found in the northern temperate and boreal regions of the earth. Spruces are large trees, from tall when mature, and can be distinguished by their whorled branches and conical...

 and Pitch Pine
Pitch Pine
The Pitch Pine, Pinus rigida, is a small-to-medium sized pine, native to eastern North America. This species occasionally hybridizes with other pine species such as Loblolly Pine , Shortleaf Pine , and Pond Pine The Pitch Pine, Pinus rigida, is a small-to-medium sized (6-30 meters or 20-100 feet)...

 stands occur as well as hickory
Hickory
Trees in the genus Carya are commonly known as hickory, derived from the Powhatan language of Virginia. The genus includes 17–19 species of deciduous trees with pinnately compound leaves and big nuts...

 and maple
Maple
Acer is a genus of trees or shrubs commonly known as maple.Maples are variously classified in a family of their own, the Aceraceae, or together with the Hippocastanaceae included in the family Sapindaceae. Modern classifications, including the Angiosperm Phylogeny Group system, favour inclusion in...

 forests.

Viburnum
Viburnum
Viburnum is a genus of about 150–175 species of shrubs or small trees in the moschatel family, Adoxaceae. Its current classification is based on molecular phylogeny...

s, winterberry, blueberry
Blueberry
Blueberries are flowering plants of the genus Vaccinium with dark-blue berries and are perennial...

, serviceberry, Virginia rose
Rosa virginiana
Rosa virginiana, commonly known as the Virginia Rose , Common Wild Rose or Prairie Rose, is a woody perennial in the rose family native to eastern North America, where it is the commonest wild rose. It is deciduous, forming a suckering shrub up to 2 metres in height, though often less. The stems...

 and male berry comprise much of the shrub understory. Other upland habitats consist of grassland
Grassland
Grasslands are areas where the vegetation is dominated by grasses and other herbaceous plants . However, sedge and rush families can also be found. Grasslands occur naturally on all continents except Antarctica...

 units and thicket units. Habitats are quite diverse, containing elements from the more southern oak-pine forests and the coniferous forests of the north. Southern Maine is where these two community types collide and blend, creating a wealth of biodiversity
Biodiversity
Biodiversity is the degree of variation of life forms within a given ecosystem, biome, or an entire planet. Biodiversity is a measure of the health of ecosystems. Biodiversity is in part a function of climate. In terrestrial habitats, tropical regions are typically rich whereas polar regions...

.

The Refuge has exemplary natural communities that include a coastal dune marsh ecosystem, Spartina saltmarsh, white oak – red oak forest, dune grassland and Pitch Pine Bog.

External links

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