Alphitonia ponderosa
Encyclopedia
Alphitonia ponderosa is a species of flowering
tree
in the buckthorn
family, Rhamnaceae
, that is endemic to the Hawaiian Islands
. It is locally known as kauila, as is the related Colubrina oppositifolia
.
are obvate, 5–15 cm (2–5.9 in) long, and have 13–25 mm (0.511811023622047–0.984251968503937 in) petiole
s. The leaves are shiny, hairless, and green on the top, but are a dull light green with rust-colored veins on the bottom.
s of A. ponderosa are polygamous
and form cymes at the bases of leaves. They are 6 mm (0.236220472440945 in) in diameter; the five sepal
s are 1.5 mm (0.0590551181102362 in) and cover five 0.75 millimetre (0.0295275590551181 in) petal
s.
of A. ponderosa is a 15 millimetre (0.590551181102362 in) diameter drupe
, which contain two to three seed
s. The seeds are shiny, oblong, and have a red covering.
, coastal mesic, and mixed mesic forests at elevations of 240 – on all main islands, but is rare except on Kauai
. It grows as a shrub
on exposed ridges.
by the Native Hawaiians
, who made laau melomelo (fishing lure
s), pāhoa (dagger
s), ihe (short spear
s), pololū (long spears), ōō (digging stick
s), hohoa (round kapa
beaters) ie kūkū (square kapa beaters), leiomano
(shark tooth
club
s), and kii
(tiki carvings) with it.
by the IUCN because of its fragmented distribution and declining population. Major threats include rat
s, pigs, deer
, competition with introduced species
of plants, and wildfire
.
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...
tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...
in the buckthorn
Buckthorn
The Buckthorns are a genus of about 100 species of shrubs or small trees from 1-10 m tall , in the buckthorn family Rhamnaceae...
family, Rhamnaceae
Rhamnaceae
Rhamnaceae, the Buckthorn family, is a large family of flowering plants, mostly trees, shrubs and some vines.The family contains 50-60 genera and approximately 870-900 species. The Rhamnaceae have a worldwide distribution, but are more common in the subtropical and tropical regions...
, that is endemic to the Hawaiian Islands
Hawaiian Islands
The Hawaiian Islands are an archipelago of eight major islands, several atolls, numerous smaller islets, and undersea seamounts in the North Pacific Ocean, extending some 1,500 miles from the island of Hawaii in the south to northernmost Kure Atoll...
. It is locally known as kauila, as is the related Colubrina oppositifolia
Colubrina oppositifolia
Colubrina oppositifolia, known as Kauila in Hawaiian, is a species of flowering tree in the buckthorn family, Rhamnaceae, that is endemic to Hawaii. It can be found in dry, coastal mesic, and mixed mesic forests at elevations of on the islands of Oahu and Hawaii . There is also one individual...
.
Description
A. ponderosa is a medium to large tree, reaching 15 – high with a trunk 20–60 cm (7.9–23.6 in) in diameter.Leaves
The alternate leavesLeaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....
are obvate, 5–15 cm (2–5.9 in) long, and have 13–25 mm (0.511811023622047–0.984251968503937 in) petiole
Petiole (botany)
In botany, the petiole is the stalk attaching the leaf blade to the stem. The petiole usually has the same internal structure as the stem. Outgrowths appearing on each side of the petiole are called stipules. Leaves lacking a petiole are called sessile, or clasping when they partly surround the...
s. The leaves are shiny, hairless, and green on the top, but are a dull light green with rust-colored veins on the bottom.
Flowers
FlowerFlower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...
s of A. ponderosa are polygamous
Plant sexuality
Plant sexuality covers the wide variety of sexual reproduction systems found across the plant kingdom. This article describes morphological aspects of sexual reproduction of plants....
and form cymes at the bases of leaves. They are 6 mm (0.236220472440945 in) in diameter; the five sepal
Sepal
A sepal is a part of the flower of angiosperms . Collectively the sepals form the calyx, which is the outermost whorl of parts that form a flower. Usually green, sepals have the typical function of protecting the petals when the flower is in bud...
s are 1.5 mm (0.0590551181102362 in) and cover five 0.75 millimetre (0.0295275590551181 in) petal
Petal
Petals are modified leaves that surround the reproductive parts of flowers. They often are brightly colored or unusually shaped to attract pollinators. Together, all of the petals of a flower are called a corolla. Petals are usually accompanied by another set of special leaves called sepals lying...
s.
Fruit
The fruitFruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...
of A. ponderosa is a 15 millimetre (0.590551181102362 in) diameter drupe
Drupe
In botany, a drupe is a fruit in which an outer fleshy part surrounds a shell of hardened endocarp with a seed inside. These fruits develop from a single carpel, and mostly from flowers with superior ovaries...
, which contain two to three seed
Seed
A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...
s. The seeds are shiny, oblong, and have a red covering.
Habitat
A. ponderosa inhabits dryHawaiian tropical dry forests
The Hawaiian tropical dry forests are a tropical dry broadleaf forest ecoregion in the Hawaiian Islands. They cover an area of on the leeward side of the main islands and the summits of Niihau and Kahoolawe. These forests are either seasonal or sclerophyllous. Annual rainfall is less than and...
, coastal mesic, and mixed mesic forests at elevations of 240 – on all main islands, but is rare except on Kauai
Kauai
Kauai or Kauai, known as Tauai in the ancient Kaua'i dialect, is geologically the oldest of the main Hawaiian Islands. With an area of , it is the fourth largest of the main islands in the Hawaiian archipelago, and the 21st largest island in the United States. Known also as the "Garden Isle",...
. It grows as a shrub
Shrub
A shrub or bush is distinguished from a tree by its multiple stems and shorter height, usually under 5–6 m tall. A large number of plants may become either shrubs or trees, depending on the growing conditions they experience...
on exposed ridges.
Uses
The reddish-brown wood of A. ponderosa is highly prized for its beauty, strength, and density. It was used as a replacement for metalMetal
A metal , is an element, compound, or alloy that is a good conductor of both electricity and heat. Metals are usually malleable and shiny, that is they reflect most of incident light...
by the Native Hawaiians
Native Hawaiians
Native Hawaiians refers to the indigenous Polynesian people of the Hawaiian Islands or their descendants. Native Hawaiians trace their ancestry back to the original Polynesian settlers of Hawaii.According to the U.S...
, who made laau melomelo (fishing lure
Fishing lure
A fishing lure is an object attached to the end of a fishing line which is designed to resemble and move like the prey of a fish. The purpose of the lure is to use movement, vibration, and colour to catch the fish's attention so it bites the hook...
s), pāhoa (dagger
Dagger
A dagger is a fighting knife with a sharp point designed or capable of being used as a thrusting or stabbing weapon. The design dates to human prehistory, and daggers have been used throughout human experience to the modern day in close combat confrontations...
s), ihe (short spear
Spear
A spear is a pole weapon consisting of a shaft, usually of wood, with a pointed head.The head may be simply the sharpened end of the shaft itself, as is the case with bamboo spears, or it may be made of a more durable material fastened to the shaft, such as flint, obsidian, iron, steel or...
s), pololū (long spears), ōō (digging stick
Digging stick
In archaeology and anthropology a digging stick is the term given to a variety of wooden implements used primarily by subsistence-based cultures to dig out underground food such as roots and tubers or burrowing animals and anthills...
s), hohoa (round kapa
Kapa
Kapa is a fabric that was made by Native Hawaiians from the bast fibres of certain species of trees and shrubs in the orders Rosales and Malvales. It is similar to tapa found elsewhere in Polynesia but differs in the methods used in its creation...
beaters) ie kūkū (square kapa beaters), leiomano
Leiomano
The leiomano is a shark-toothed club used by various Polynesian tribes, but mostly by the native Hawaiians.Leiomano is a word in the Hawaiian language and may have been derived from lei o manō, which means "a shark's lei."...
(shark tooth
Shark tooth
A shark tooth is one of the numerous teeth of a shark. Sharks continually shed their teeth, and some Carcharhiniformes shed approximately 35,000 teeth in a lifetime. In some geological formations, shark's teeth are a common fossil...
club
Club (weapon)
A club is among the simplest of all weapons. A club is essentially a short staff, or stick, usually made of wood, and wielded as a weapon since prehistoric times....
s), and kii
Tiki
Tiki refers to large wood and stone carvings of humanoid forms in Central Eastern Polynesian cultures of the Pacific Ocean. The term is also used in Māori mythology where Tiki is the first man, created by either Tūmatauenga or Tāne. He found the first woman, Marikoriko, in a pond – she seduced him...
(tiki carvings) with it.
Conservation
A. ponderosa is considered a vulnerable speciesVulnerable species
On 30 January 2010, the IUCN Red List of Threatened Species identified 9694 Vulnerable species, subspecies and varieties, stocks and sub-populations.-References:...
by the IUCN because of its fragmented distribution and declining population. Major threats include rat
Rat
Rats are various medium-sized, long-tailed rodents of the superfamily Muroidea. "True rats" are members of the genus Rattus, the most important of which to humans are the black rat, Rattus rattus, and the brown rat, Rattus norvegicus...
s, pigs, deer
Deer
Deer are the ruminant mammals forming the family Cervidae. Species in the Cervidae family include white-tailed deer, elk, moose, red deer, reindeer, fallow deer, roe deer and chital. Male deer of all species and female reindeer grow and shed new antlers each year...
, competition with introduced species
Invasive species
"Invasive species", or invasive exotics, is a nomenclature term and categorization phrase used for flora and fauna, and for specific restoration-preservation processes in native habitats, with several definitions....
of plants, and wildfire
Wildfire
A wildfire is any uncontrolled fire in combustible vegetation that occurs in the countryside or a wilderness area. Other names such as brush fire, bushfire, forest fire, desert fire, grass fire, hill fire, squirrel fire, vegetation fire, veldfire, and wilkjjofire may be used to describe the same...
.