Diphasiastrum
Encyclopedia
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...
of clubmosses in the plant family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...
Lycopodiaceae
Lycopodiaceae
The Lycopodiaceae is a family of primitive vascular plants, including all of the core clubmosses. These plants bear spores on specialized structures at the apex of a shoot; they resemble a tiny battle club, from which the common name derives...
. It is closely related to the genus Lycopodium
Lycopodium
Lycopodium is a genus of clubmosses, also known as ground pines or creeping cedar, in the family Lycopodiaceae, a family of fern-allies...
, and some botanists treat it within a broad view of that genus as a section, Lycopodium sect. Complanata. Some species superficially resemble diminutive gymnosperm
Gymnosperm
The gymnosperms are a group of seed-bearing plants that includes conifers, cycads, Ginkgo, and Gnetales. The term "gymnosperm" comes from the Greek word gymnospermos , meaning "naked seeds", after the unenclosed condition of their seeds...
s and have been given common names such as ground-pine or ground-cedar.
The genus has a subcosmopolitan
Cosmopolitan distribution
In biogeography, a taxon is said to have a cosmopolitan distribution if its range extends across all or most of the world in appropriate habitats. For instance, the killer whale has a cosmopolitan distribution, extending over most of the world's oceans. Other examples include humans, the lichen...
distribution, in much of the Northern Hemisphere
Northern Hemisphere
The Northern Hemisphere is the half of a planet that is north of its equator—the word hemisphere literally means “half sphere”. It is also that half of the celestial sphere north of the celestial equator...
, south in mountain
Mountain
Image:Himalaya_annotated.jpg|thumb|right|The Himalayan mountain range with Mount Everestrect 58 14 160 49 Chomo Lonzorect 200 28 335 52 Makalurect 378 24 566 45 Mount Everestrect 188 581 920 656 Tibetan Plateaurect 250 406 340 427 Rong River...
s to South America
South America
South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...
(reaching furthest south in Jujuy Province
Jujuy Province
Jujuy is a province of Argentina, located in the extreme northwest of the country, at the borders with Chile and Bolivia. The only neighboring Argentine province is Salta to the east and south.-History:...
, northwest Argentina
Argentina
Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...
), New Guinea
New Guinea
New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...
and the Marquesas Islands
Marquesas Islands
The Marquesas Islands enana and Te Fenua `Enata , both meaning "The Land of Men") are a group of volcanic islands in French Polynesia, an overseas collectivity of France in the southern Pacific Ocean. The Marquesas are located at 9° 00S, 139° 30W...
in the Pacific Ocean
Pacific Ocean
The Pacific Ocean is the largest of the Earth's oceanic divisions. It extends from the Arctic in the north to the Southern Ocean in the south, bounded by Asia and Australia in the west, and the Americas in the east.At 165.2 million square kilometres in area, this largest division of the World...
, but confined to climate
Climate
Climate encompasses the statistics of temperature, humidity, atmospheric pressure, wind, rainfall, atmospheric particle count and other meteorological elemental measurements in a given region over long periods...
s with high humidity
Humidity
Humidity is a term for the amount of water vapor in the air, and can refer to any one of several measurements of humidity. Formally, humid air is not "moist air" but a mixture of water vapor and other constituents of air, and humidity is defined in terms of the water content of this mixture,...
for most or all of the year (or, in cool climates, protected by snow
Snow
Snow is a form of precipitation within the Earth's atmosphere in the form of crystalline water ice, consisting of a multitude of snowflakes that fall from clouds. Since snow is composed of small ice particles, it is a granular material. It has an open and therefore soft structure, unless packed by...
cover in winter).
There are 16 species, and numerous natural hybrids in the genus; many of the hybrids are fertile, allowing their occurrence to become frequent, sometimes more so than the parent species. The basal chromosome count for this genus is n=23, which is distinctively different than other lycopods.
Several species have been used economically for their spores, which are harvested as Lycopodium powder
Lycopodium powder
Lycopodium powder is a yellow-tan dust-like powder historically used as a flash powder. It is composed of the dry spores of clubmoss plants, various fern relatives principally in the genera Lycopodium and Diphasiastrum...
.
Species
- Diphasiastrum alpinumDiphasiastrum alpinumDiphasiastrum alpinum, Alpine Clubmoss, is a species of clubmoss. It is found in mountains and moors often with Calluna and grasses....
Alpine Clubmoss (circumpolar, subarctic and alpine) - Diphasiastrum angustiramosum (New GuineaNew GuineaNew Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...
) - Diphasiastrum complanatumDiphasiastrum complanatumDiphasiastrum complanatum Holub is a species of clubmoss native to dry coniferous forests throughout the Holarctic Kingdom...
Flat-stemmed Clubmoss, Northern Running-pine, or Ground Cedar (circumpolar, cool temperate) - Diphasiastrum digitatumDiphasiastrum digitatumDiphasiastrum digitatum is most commonly known as Groundcedar, along with other members of its genus, but the common name Fan Clubmoss can be used to refer to it specifically. It is the most common species of Diphasiastrum in North America. It is a type of plant known as a clubmoss, which is...
Fan Clubmoss, Southern Running-pine, or Running Cedar (eastern CanadaCanadaCanada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...
, northeastern United StatesUnited StatesThe United States of America is a federal constitutional republic comprising fifty states and a federal district...
) - Diphasiastrum fawcettii (JamaicaJamaicaJamaica is an island nation of the Greater Antilles, in length, up to in width and 10,990 square kilometres in area. It is situated in the Caribbean Sea, about south of Cuba, and west of Hispaniola, the island harbouring the nation-states Haiti and the Dominican Republic...
, HispaniolaHispaniolaHispaniola is a major island in the Caribbean, containing the two sovereign states of the Dominican Republic and Haiti. The island is located between the islands of Cuba to the west and Puerto Rico to the east, within the hurricane belt...
) - Diphasiastrum henryanum (Marquesas IslandsMarquesas IslandsThe Marquesas Islands enana and Te Fenua `Enata , both meaning "The Land of Men") are a group of volcanic islands in French Polynesia, an overseas collectivity of France in the southern Pacific Ocean. The Marquesas are located at 9° 00S, 139° 30W...
) - Diphasiastrum madeirense (MadeiraMadeiraMadeira is a Portuguese archipelago that lies between and , just under 400 km north of Tenerife, Canary Islands, in the north Atlantic Ocean and an outermost region of the European Union...
, AzoresAzoresThe Archipelago of the Azores is composed of nine volcanic islands situated in the middle of the North Atlantic Ocean, and is located about west from Lisbon and about east from the east coast of North America. The islands, and their economic exclusion zone, form the Autonomous Region of the...
) - Diphasiastrum montellii (RussiaRussiaRussia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...
; syn. D. complanatum subsp. montellii) - Diphasiastrum multispicatum (TaiwanTaiwanTaiwan , also known, especially in the past, as Formosa , is the largest island of the same-named island group of East Asia in the western Pacific Ocean and located off the southeastern coast of mainland China. The island forms over 99% of the current territory of the Republic of China following...
, PhilippinesPhilippinesThe Philippines , officially known as the Republic of the Philippines , is a country in Southeast Asia in the western Pacific Ocean. To its north across the Luzon Strait lies Taiwan. West across the South China Sea sits Vietnam...
) - Diphasiastrum nikoense (JapanJapanJapan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...
; syn. D. sitchense var. nikoense) - Diphasiastrum platyrhizoma (BorneoBorneoBorneo is the third largest island in the world and is located north of Java Island, Indonesia, at the geographic centre of Maritime Southeast Asia....
, SumatraSumatraSumatra is an island in western Indonesia, westernmost of the Sunda Islands. It is the largest island entirely in Indonesia , and the sixth largest island in the world at 473,481 km2 with a population of 50,365,538...
) - Diphasiastrum sitchense Alaskan Clubmoss (northern North AmericaNorth AmericaNorth America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...
) - Diphasiastrum thyoides (CaribbeanCaribbeanThe Caribbean is a crescent-shaped group of islands more than 2,000 miles long separating the Gulf of Mexico and the Caribbean Sea, to the west and south, from the Atlantic Ocean, to the east and north...
, CentralCentral AmericaCentral America is the central geographic region of the Americas. It is the southernmost, isthmian portion of the North American continent, which connects with South America on the southeast. When considered part of the unified continental model, it is considered a subcontinent...
and South AmericaSouth AmericaSouth America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...
) - Diphasiastrum tristachyum Blue Clubmoss, Blue Ground-cedar (circumpolar, cool temperate; syn. D. complanatum subsp. chamaecyparissus)
- Diphasiastrum veitchii Veitch's Clubmoss (eastern Himalaya east to TaiwanTaiwanTaiwan , also known, especially in the past, as Formosa , is the largest island of the same-named island group of East Asia in the western Pacific Ocean and located off the southeastern coast of mainland China. The island forms over 99% of the current territory of the Republic of China following...
) - Diphasiastrum wightianum (southeast AsiaAsiaAsia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...
, New GuineaNew GuineaNew Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...
)
Selected hybrids
- Diphasiastrum × habereri (D . digitatum × D. tristachyum)
- Diphasiastrum × issleri (D. alpinum × D. complanatum)
- Diphasiastrum × sabinifolium (D. sitchense × D. tristachyum)
- Diphasiastrum × zeilleri (D. complanatum × D. tristachyum)