Pandalus borealis
Encyclopedia
Pandalus borealis is a species of shrimp
found in cold parts of the Atlantic
and Pacific Ocean
s. Many different common name
s are used, including pink shrimp, deepwater prawn, deep-sea prawn, great northern prawn, and northern shrimp.
, Canada's
eastern seaboard (off Newfoundland and Labrador
and eastern Baffin Island
in Nunavut
), southern and eastern Greenland
, Iceland
, Svalbard
, Norway
and the North Sea
as far south as the English Channel
. In the Pacific, P. b. eous is found from Japan
, through the Sea of Okhotsk
, across the Bering Strait
, and as far south in North America
as Washington state.
The shrimp are hermaphroditic
. They start out male, but after year or two, their testicles turn to ovaries and they complete their lives as females.
since the early 1900s in Norway
, and later in other countries following Johan Hjort's
practical discoveries of how to locate them. In Canada, these shrimp are sold peeled, cooked and frozen in bags in supermarkets, and are consumed as appetizers.
Shrimp alkaline phosphatase
(SAP), an enzyme
used in molecular biology
, is obtained from Pandalus borealis. The species' carapace
is a source of chitosan
, a versatile chemical used for such different applications as treating bleeding wounds, filtering wine or improving the soil in organic farming
.
In Canada, the annual harvest limit is set to 164,000 tonnes (2008). The Canadian fishery began in the 1980s and expanded in 1990s.
Shrimp
Shrimp are swimming, decapod crustaceans classified in the infraorder Caridea, found widely around the world in both fresh and salt water. Adult shrimp are filter feeding benthic animals living close to the bottom. They can live in schools and can swim rapidly backwards. Shrimp are an important...
found in cold parts of the Atlantic
Atlantic Ocean
The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...
and Pacific Ocean
Pacific Ocean
The Pacific Ocean is the largest of the Earth's oceanic divisions. It extends from the Arctic in the north to the Southern Ocean in the south, bounded by Asia and Australia in the west, and the Americas in the east.At 165.2 million square kilometres in area, this largest division of the World...
s. Many different common name
Common name
A common name of a taxon or organism is a name in general use within a community; it is often contrasted with the scientific name for the same organism...
s are used, including pink shrimp, deepwater prawn, deep-sea prawn, great northern prawn, and northern shrimp.
Distribution
P. borealis lives at depths of 20–1330 m (65.6–4,363.5 ft), usually on soft muddy bottoms, in waters with a temperature of 2–14 °C (35.6–57.2 F). The distribution of the nominate subspecies P. b. borealis in the Atlantic ranges from New EnglandNew England
New England is a region in the northeastern corner of the United States consisting of the six states of Maine, New Hampshire, Vermont, Massachusetts, Rhode Island, and Connecticut...
, Canada's
Canada
Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...
eastern seaboard (off Newfoundland and Labrador
Newfoundland and Labrador
Newfoundland and Labrador is the easternmost province of Canada. Situated in the country's Atlantic region, it incorporates the island of Newfoundland and mainland Labrador with a combined area of . As of April 2011, the province's estimated population is 508,400...
and eastern Baffin Island
Baffin Island
Baffin Island in the Canadian territory of Nunavut is the largest island in the Canadian Arctic Archipelago, the largest island in Canada and the fifth largest island in the world. Its area is and its population is about 11,000...
in Nunavut
Nunavut
Nunavut is the largest and newest federal territory of Canada; it was separated officially from the Northwest Territories on April 1, 1999, via the Nunavut Act and the Nunavut Land Claims Agreement Act, though the actual boundaries had been established in 1993...
), southern and eastern Greenland
Greenland
Greenland is an autonomous country within the Kingdom of Denmark, located between the Arctic and Atlantic Oceans, east of the Canadian Arctic Archipelago. Though physiographically a part of the continent of North America, Greenland has been politically and culturally associated with Europe for...
, Iceland
Iceland
Iceland , described as the Republic of Iceland, is a Nordic and European island country in the North Atlantic Ocean, on the Mid-Atlantic Ridge. Iceland also refers to the main island of the country, which contains almost all the population and almost all the land area. The country has a population...
, Svalbard
Svalbard
Svalbard is an archipelago in the Arctic, constituting the northernmost part of Norway. It is located north of mainland Europe, midway between mainland Norway and the North Pole. The group of islands range from 74° to 81° north latitude , and from 10° to 35° east longitude. Spitsbergen is the...
, Norway
Norway
Norway , officially the Kingdom of Norway, is a Nordic unitary constitutional monarchy whose territory comprises the western portion of the Scandinavian Peninsula, Jan Mayen, and the Arctic archipelago of Svalbard and Bouvet Island. Norway has a total area of and a population of about 4.9 million...
and the North Sea
North Sea
In the southwest, beyond the Straits of Dover, the North Sea becomes the English Channel connecting to the Atlantic Ocean. In the east, it connects to the Baltic Sea via the Skagerrak and Kattegat, narrow straits that separate Denmark from Norway and Sweden respectively...
as far south as the English Channel
English Channel
The English Channel , often referred to simply as the Channel, is an arm of the Atlantic Ocean that separates southern England from northern France, and joins the North Sea to the Atlantic. It is about long and varies in width from at its widest to in the Strait of Dover...
. In the Pacific, P. b. eous is found from Japan
Japan
Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...
, through the Sea of Okhotsk
Sea of Okhotsk
The Sea of Okhotsk is a marginal sea of the western Pacific Ocean, lying between the Kamchatka Peninsula on the east, the Kuril Islands on the southeast, the island of Hokkaidō to the far south, the island of Sakhalin along the west, and a long stretch of eastern Siberian coast along the west and...
, across the Bering Strait
Bering Strait
The Bering Strait , known to natives as Imakpik, is a sea strait between Cape Dezhnev, Chukotka Autonomous Okrug, Russia, the easternmost point of the Asian continent and Cape Prince of Wales, Alaska, USA, the westernmost point of the North American continent, with latitude of about 65°40'N,...
, and as far south in North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...
as Washington state.
Physiology
In their 8 year lifespan, males can reach a length of 120 mm (4.7 in), while females can reach 165 mm (6.5 in) long.The shrimp are hermaphroditic
Hermaphrodite
In biology, a hermaphrodite is an organism that has reproductive organs normally associated with both male and female sexes.Many taxonomic groups of animals do not have separate sexes. In these groups, hermaphroditism is a normal condition, enabling a form of sexual reproduction in which both...
. They start out male, but after year or two, their testicles turn to ovaries and they complete their lives as females.
Uses by humans
P. borealis is an important food resource, and has been widely fishedShrimp fishery
A shrimp fishery is a fishery directed toward harvesting either shrimp or prawns. .-Commercial shrimping:...
since the early 1900s in Norway
Norway
Norway , officially the Kingdom of Norway, is a Nordic unitary constitutional monarchy whose territory comprises the western portion of the Scandinavian Peninsula, Jan Mayen, and the Arctic archipelago of Svalbard and Bouvet Island. Norway has a total area of and a population of about 4.9 million...
, and later in other countries following Johan Hjort's
Johan Hjort
Johan Hjort FRS was a Norwegian fisheries scientist, marine zoologist, and oceanographer. He was among the most prominent and influential marine zoologists of his time.- The early years :...
practical discoveries of how to locate them. In Canada, these shrimp are sold peeled, cooked and frozen in bags in supermarkets, and are consumed as appetizers.
Shrimp alkaline phosphatase
Alkaline phosphatase
Alkaline phosphatase is a hydrolase enzyme responsible for removing phosphate groups from many types of molecules, including nucleotides, proteins, and alkaloids. The process of removing the phosphate group is called dephosphorylation...
(SAP), an enzyme
Enzyme
Enzymes are proteins that catalyze chemical reactions. In enzymatic reactions, the molecules at the beginning of the process, called substrates, are converted into different molecules, called products. Almost all chemical reactions in a biological cell need enzymes in order to occur at rates...
used in molecular biology
Molecular biology
Molecular biology is the branch of biology that deals with the molecular basis of biological activity. This field overlaps with other areas of biology and chemistry, particularly genetics and biochemistry...
, is obtained from Pandalus borealis. The species' carapace
Carapace
A carapace is a dorsal section of the exoskeleton or shell in a number of animal groups, including arthropods such as crustaceans and arachnids, as well as vertebrates such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron.-Crustaceans:In crustaceans, the...
is a source of chitosan
Chitosan
Chitosan is a linear polysaccharide composed of randomly distributed β--linked D-glucosamine and N-acetyl-D-glucosamine...
, a versatile chemical used for such different applications as treating bleeding wounds, filtering wine or improving the soil in organic farming
Organic farming
Organic farming is the form of agriculture that relies on techniques such as crop rotation, green manure, compost and biological pest control to maintain soil productivity and control pests on a farm...
.
Management
Northern shrimp have a short life, which contributes to a variable stock on a yearly basis. However, the species is not considered overfished due to a large amount reported and a large amount harvested.In Canada, the annual harvest limit is set to 164,000 tonnes (2008). The Canadian fishery began in the 1980s and expanded in 1990s.