Groundsel
Encyclopedia
Senecio vulgaris, often known by the common name Common groundsel, a humble in appearance member of the Asteraceae
family
and Senecio
genus
, is a tenacious deciduous
annual
whose presence now encompasses the globe in a wide area of easy to somewhat difficult growing conditions.
The discussion of Common groundsel dates back to the 1st century and more recently, it is the subject of much contradictory and reactionary information about where it came from, how it got there, whether is it really hurting the crops, how to get rid of it, and how dangerous it is when ingested by various animals.
Leaves of Senecio vulgaris grow directly from the stem, sessile or lacking their own stem (petiole), alternating in direction along the length of the plant, two rounded lobes at the base of the stem (auriculate) and sub-clasping above. Leaves are pinnately lobed and +2.4 inches (61 mm) long and 1 inches (25.4 mm) wide and get smaller as up the plant. Leaves are covered sparsely with soft, smooth, fine hairs. Lobes typically sharp to rounded saw-toothed.
The hollow
succulent stems branch at the tops and from the base.
Stems and leaves can both host the Cinerarea leaf rust
.
Open clusters of 8 to 10 small cylinder shaped rayless yellow
flower heads ¼ to ½ inch (6 to 13 mm) with a highly conspicuous ring of black tipped bract
s at the base of the inflorescence as is characteristic of many members of the genus Senecio
.
The name for the genus Senecio is probably derived from Senex (an old man), in reference to its downy head of seeds; "the flower of this herb hath white hair and when the wind bloweth it away, then it appeareth like a bald-headed man" and like its family, flowers of Senecio vulgaris are succeeded by downy globed heads of seed. The seeds
are achene
, include a pappus
and become sticky when wet.
Laboratory tests have suggested maximum seed scattering distances of 2.1 and 3.2 yd (1.9 and 2.9 m) at wind speeds of 6.8 and 10.2 mph (10.9 and 16.4 km/h) respectively (affected by plant height) suggests that it was more than wind that spread these groundsel seeds throughout the world.
The average weight of 1000 seeds is 0.21 gram (2,200,000 seeds per pound) and experienced a 100% germination
success before drying and storage and an 87% germination success after drying and 3 years of cool dry storage.
In simple models for seed emergence prediction, soil thermal time
did not predict the timing and extent of seedling
emergence as well as hydrothermal time
(warm rain).
The root system consists of a shallow taproot. This plant spreads by reseeding itself.
Groundsel acts as a host for the fungus that causes black root rot
in peas
, alfalfa
, soybeans, carrot
s, tomato
es, red clover
, peanut
s, cucurbit
s, cotton
, citrus
, chickpea
s, and several ornamental flowering plants; a list of flowering plant
s that can host their own fungus as well.
deciduous annual plant that grows willingly in disturbed sites, waste places, roadsides, gardens, nurseries, orchards, vineyards, landscaped areas, agricultural lands,
at altitudes up to 1600 feet (487.7 m)
and is, additionally, self-pollinating
producing 1,700 seeds per plant with three generations per year.
Seeds are dispersed by wind and also cling to clothing and animal fur, and as contaminates of commercially exchanged seeds; the relocation of this plant throughout the planet
has been difficult if not impossible to contain.
There is conflicting information about the native status of Senecio vulgaris in various locations. The United States Department of Agriculture (USDA)
, Natural Resources Conservation Service Plants Profile Database
considers it to be native to all 50 of the United States of America
, Canada
, Greenland
, Saint Pierre and Miquelon, the same USDA through the Germplasm Resources Information Network (GRIN)
considers it to be native only to parts of Afro-Eurasia
. The Integrated Taxonomic Information System Organization (ITIS)
, a partnership between many United States federal government departments and agencies
states that the species has been introduced to the 50 United States,
and the online journal Flora of North America
calls it "probably introduced" to areas north of Mexico.
Individual research groups claim it is not native to areas they oversee: Florida
,
Washington,
Wisconsin
,
Saskatchewan
,
British Columbia
,
Missouri
.
The United States Geological Survey
reports that Common Groundsel is exotic to all 50 states and all Canadian provinces with the exception of Georgia
, Kentucky
, Massachusetts
, and Labrador
.
Native
Current
for Canaries
and Finches
and it is available all year
round.
Senecio vulgaris seed has been found in the droppings of sparrow
s, and seedlings have been raised from the excreta of various birds. Seed has also been found in cow manure
.
Some Lepidoptera
species eat many of the Senecio genus; additional studies via electrophysiological recordings
have shown that the taste sensilla
of the Cinnabar moth
larvae
respond (get excited) specifically to the pyrrolizidine alkaloids, which all Senecio are known to contain.
Moths and caterpillars
The Senecio genus also are host to other insects:
Beetles
Flies
and other insects that are not listed here.
The ragwort flea beetle and ragwort seed fly have been approved and released for Senecio control in California
, Australia and elsewhere.
Fungus
Most Senecio, including S. squalidus are susceptible to rust and other fungus
and mildews
:
and other fungus that are not listed here.
being both non-indigenous
to most if not all of the Americas and having a reputation for being hepatotoxic
to livestock
and to humans.
As a plant that is reported to be both poison
ous for human ingestion
and also medicinal; much of the contradiction can be found by closely reviewing the words that are used and the dose (amount) of the poisonous substance that is ingested to prove either claim. All species of the genus Senecio contain pyrrolizidine alkaloids
(e.g., senecionine
) a substance that when a human has chronic exposure
can cause irreversible liver damage.
Common groundsel as a medicinal herb does not seem to be recommended very often since 1931, when it was recommended as a diaphoretic
, an antiscorbutic
, a purgative
, a diuretic
and an anthelmintic
, which was a demotion as it was previously suggested for the expelling of gravel of the kidneys and reins
by Pedanius Dioscorides
in the 70s-90s, for use as poultice
s by John Gerard
in the late 16th century and as a cure for epilepsy
by Nicholas Culpeper
in the 17th century. More current information is contradictory about the dangers of the ingestion of Groundsel. A heavily referenced paper from 1989 suggests that the response is immediate and gives pre-ambulatory care recommendations. A Canadian poisonous plants information database references a paper from 1990 in presenting this prenatal warning: "In a case of prenatal exposure, a mother ingested tea containing an estimated 0.343 milligram of senecionine, resulting in fatal veno-occlusive disease in a newborn infant." Information about the pyrrolizidine alkaloids, the substance present in Senecio vulgaris is much less contradictory and all warn of accumulation of the alkaloid.
Certain pyrrolizidine alkaloids are non-toxic precursors that are converted to toxic metabolite
s in the body in a process called toxification
Livestock
Linnaeus
is cited to have claimed that "goat
s and swine
eat this common plant freely, cows
being not partial to it and horse
s and sheep
declining to touch it, but not only are caged birds
fond of it (the seeds), but its leaves and seeds afford food for many of our wild species (rabbit
s were given as an example)." More recent studies claim that the lethal
amount that cattle or horses need to consume is 7% of their body weight
(example: 50 pounds (23 kg) would need to be consumed by a cow weighing 700 pounds (318 kg)). Lesser amounts cause the liver
to lose function but is not apparent until the animal is stressed (by new feed or location, pregnancy, a different toxin
, etc.). Sheep and goats have rumen
bacteria
that detoxify the alkaloid
s and are able to consume twice their body weight of this and other species of genus Senecio. The alkaloids responsible are not destroyed by drying or by fermentation in silage.
or with crops
. Senecio vulgaris is not known to be a strong competitor but it has been known to reduce mint production.
There is evidence that it is not a strong invasive and sometimes protective of critically endangered native plants.
Control
The approximately 22 millimetre (0.866141732283465 in) long
pappus
seeds of Senecio vulgaris, each plant capable of producing 25,000 or more seeds (1,700 seeds per plant are more likely) with three generations of the plant per year; seeds that are widely dispersed by the wind,
have been identified as a contaminant of cereal and vegetable seeds and a poison to some livestock; there is some inspiration to understand the growth stages and determine some control methods.
Asteraceae
The Asteraceae or Compositae , is an exceedingly large and widespread family of vascular plants. The group has more than 22,750 currently accepted species, spread across 1620 genera and 12 subfamilies...
family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...
and Senecio
Senecio
Senecio is a genus of the daisy family that includes ragworts and groundsels. The flower heads are normally rayed, completely yellow, and the heads are borne in branched clusters...
genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...
, is a tenacious deciduous
Deciduous
Deciduous means "falling off at maturity" or "tending to fall off", and is typically used in reference to trees or shrubs that lose their leaves seasonally, and to the shedding of other plant structures such as petals after flowering or fruit when ripe...
annual
Annual plant
An annual plant is a plant that usually germinates, flowers, and dies in a year or season. True annuals will only live longer than a year if they are prevented from setting seed...
whose presence now encompasses the globe in a wide area of easy to somewhat difficult growing conditions.
The discussion of Common groundsel dates back to the 1st century and more recently, it is the subject of much contradictory and reactionary information about where it came from, how it got there, whether is it really hurting the crops, how to get rid of it, and how dangerous it is when ingested by various animals.
Description
Standing only between 4 and 16 inches (10 to 41 cm) tall, bright florets mostly hidden by the characteristic bract giving it the appearance of never opening flowers and with a life span of 5–6 weeks, the self fertilizing Senecio vulgaris lives humbly among and occasionally under the other weeds and is easy not to notice.- Leaves and stems
Leaves of Senecio vulgaris grow directly from the stem, sessile or lacking their own stem (petiole), alternating in direction along the length of the plant, two rounded lobes at the base of the stem (auriculate) and sub-clasping above. Leaves are pinnately lobed and +2.4 inches (61 mm) long and 1 inches (25.4 mm) wide and get smaller as up the plant. Leaves are covered sparsely with soft, smooth, fine hairs. Lobes typically sharp to rounded saw-toothed.
The hollow
succulent stems branch at the tops and from the base.
Stems and leaves can both host the Cinerarea leaf rust
Coleosporium tussilaginis
Coleosporium tussilaginis is species of rust fungus in the Coleosporiaceae family. It is a plant pathogen....
.
- Flowers
Open clusters of 8 to 10 small cylinder shaped rayless yellow
Yellow
Yellow is the color evoked by light that stimulates both the L and M cone cells of the retina about equally, with no significant stimulation of the S cone cells. Light with a wavelength of 570–590 nm is yellow, as is light with a suitable mixture of red and green...
flower heads ¼ to ½ inch (6 to 13 mm) with a highly conspicuous ring of black tipped bract
Bract
In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis, or cone scale. Bracts are often different from foliage leaves. They may be smaller, larger, or of a different color, shape, or texture...
s at the base of the inflorescence as is characteristic of many members of the genus Senecio
Senecio
Senecio is a genus of the daisy family that includes ragworts and groundsels. The flower heads are normally rayed, completely yellow, and the heads are borne in branched clusters...
.
- Seeds
The name for the genus Senecio is probably derived from Senex (an old man), in reference to its downy head of seeds; "the flower of this herb hath white hair and when the wind bloweth it away, then it appeareth like a bald-headed man" and like its family, flowers of Senecio vulgaris are succeeded by downy globed heads of seed. The seeds
Spore
In biology, a spore is a reproductive structure that is adapted for dispersal and surviving for extended periods of time in unfavorable conditions. Spores form part of the life cycles of many bacteria, plants, algae, fungi and some protozoa. According to scientist Dr...
are achene
Achene
An achene is a type of simple dry fruit produced by many species of flowering plants. Achenes are monocarpellate and indehiscent...
, include a pappus
Pappus (flower structure)
The pappus is the modified calyx, the part of an individual disk, ray or ligule floret surrounding the base of the corolla, in flower heads of the plant family Asteraceae. The pappus may be composed of bristles , awns, scales, or may be absent. In some species, the pappus is too small to see...
and become sticky when wet.
Laboratory tests have suggested maximum seed scattering distances of 2.1 and 3.2 yd (1.9 and 2.9 m) at wind speeds of 6.8 and 10.2 mph (10.9 and 16.4 km/h) respectively (affected by plant height) suggests that it was more than wind that spread these groundsel seeds throughout the world.
The average weight of 1000 seeds is 0.21 gram (2,200,000 seeds per pound) and experienced a 100% germination
Germination
Germination is the process in which a plant or fungus emerges from a seed or spore, respectively, and begins growth. The most common example of germination is the sprouting of a seedling from a seed of an angiosperm or gymnosperm. However the growth of a sporeling from a spore, for example the...
success before drying and storage and an 87% germination success after drying and 3 years of cool dry storage.
In simple models for seed emergence prediction, soil thermal time
Soil thermal properties
The thermal properties of soil are a component of soil physics that has found important uses in engineering, climatology and agriculture. These properties influence how energy is partitioned in the soil profile...
did not predict the timing and extent of seedling
Seedling
thumb|Monocot and dicot seedlingsA seedling is a young plant sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle , the hypocotyl , and the cotyledons...
emergence as well as hydrothermal time
(warm rain).
- Roots
The root system consists of a shallow taproot. This plant spreads by reseeding itself.
Groundsel acts as a host for the fungus that causes black root rot
Thielaviopsis basicola
Thielaviopsis basicola is a plant pathogen.- External links :* *...
in peas
PEAS
P.E.A.S. is an acronym in artificial intelligence that stands for Performance, Environment, Actuators, Sensors.-Performance:Performance is a function that measures the quality of the actions the agent did....
, alfalfa
Alfalfa
Alfalfa is a flowering plant in the pea family Fabaceae cultivated as an important forage crop in the US, Canada, Argentina, France, Australia, the Middle East, South Africa, and many other countries. It is known as lucerne in the UK, France, Australia, South Africa and New Zealand, and known as...
, soybeans, carrot
Carrot
The carrot is a root vegetable, usually orange in colour, though purple, red, white, and yellow varieties exist. It has a crisp texture when fresh...
s, tomato
Tomato
The word "tomato" may refer to the plant or the edible, typically red, fruit which it bears. Originating in South America, the tomato was spread around the world following the Spanish colonization of the Americas, and its many varieties are now widely grown, often in greenhouses in cooler...
es, red clover
Red clover
Trifolium pratense is a species of clover, native to Europe, Western Asia and northwest Africa, but planted and naturalised in many other regions....
, peanut
Peanut
The peanut, or groundnut , is a species in the legume or "bean" family , so it is not a nut. The peanut was probably first cultivated in the valleys of Peru. It is an annual herbaceous plant growing tall...
s, cucurbit
Cucurbit
Cucurbit may refer to:*A plant of the family Cucurbitaceae*The lower part of an alembic*Cucurbit flute, Chinese musical instrument...
s, cotton
Cotton
Cotton is a soft, fluffy staple fiber that grows in a boll, or protective capsule, around the seeds of cotton plants of the genus Gossypium. The fiber is almost pure cellulose. The botanical purpose of cotton fiber is to aid in seed dispersal....
, citrus
Citrus
Citrus is a common term and genus of flowering plants in the rue family, Rutaceae. Citrus is believed to have originated in the part of Southeast Asia bordered by Northeastern India, Myanmar and the Yunnan province of China...
, chickpea
Chickpea
The chickpea is a legume of the family Fabaceae, subfamily Faboideae...
s, and several ornamental flowering plants; a list of flowering plant
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...
s that can host their own fungus as well.
Common names
.Distribution
Senecio vulgaris is a frost resistantdeciduous annual plant that grows willingly in disturbed sites, waste places, roadsides, gardens, nurseries, orchards, vineyards, landscaped areas, agricultural lands,
at altitudes up to 1600 feet (487.7 m)
and is, additionally, self-pollinating
producing 1,700 seeds per plant with three generations per year.
Seeds are dispersed by wind and also cling to clothing and animal fur, and as contaminates of commercially exchanged seeds; the relocation of this plant throughout the planet
Earth
Earth is the third planet from the Sun, and the densest and fifth-largest of the eight planets in the Solar System. It is also the largest of the Solar System's four terrestrial planets...
has been difficult if not impossible to contain.
There is conflicting information about the native status of Senecio vulgaris in various locations. The United States Department of Agriculture (USDA)
United States Department of Agriculture
The United States Department of Agriculture is the United States federal executive department responsible for developing and executing U.S. federal government policy on farming, agriculture, and food...
, Natural Resources Conservation Service Plants Profile Database
Natural Resources Conservation Service
The Natural Resources Conservation Service , formerly known as the Soil Conservation Service , is an agency of the United States Department of Agriculture that provides technical assistance to farmers and other private landowners and managers.Its name was changed in 1994 during the Presidency of...
considers it to be native to all 50 of the United States of America
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...
, Canada
Canada
Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...
, Greenland
Greenland
Greenland is an autonomous country within the Kingdom of Denmark, located between the Arctic and Atlantic Oceans, east of the Canadian Arctic Archipelago. Though physiographically a part of the continent of North America, Greenland has been politically and culturally associated with Europe for...
, Saint Pierre and Miquelon, the same USDA through the Germplasm Resources Information Network (GRIN)
Germplasm Resources Information Network
Germplasm Resources Information Network or GRIN is an online software project of National Genetic Resources Program of USDA to provide germplasm information about plants, animals, microbes and invertebrates.-Sub-Projects:...
considers it to be native only to parts of Afro-Eurasia
Afro-Eurasia
Afro-Eurasia or less commonly Afrasia or Eurafrasia is the term used to describe the largest landmass on earth. It may be defined as a supercontinent, consisting of Africa and Eurasia...
. The Integrated Taxonomic Information System Organization (ITIS)
Integrated Taxonomic Information System
The Integrated Taxonomic Information System is a partnership designed to provide consistent and reliable information on the taxonomy of biological species. ITIS was originally formed in 1996 as an interagency group within the U.S...
, a partnership between many United States federal government departments and agencies
Federal government of the United States
The federal government of the United States is the national government of the constitutional republic of fifty states that is the United States of America. The federal government comprises three distinct branches of government: a legislative, an executive and a judiciary. These branches and...
states that the species has been introduced to the 50 United States,
and the online journal Flora of North America
Flora of North America
The Flora of North America North of Mexico is a multivolume work describing the native plants of North America. These days much of the Flora is available . The work is expected to fill 30 volumes when completed...
calls it "probably introduced" to areas north of Mexico.
Individual research groups claim it is not native to areas they oversee: Florida
Florida
Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...
,
Washington,
Wisconsin
Wisconsin
Wisconsin is a U.S. state located in the north-central United States and is part of the Midwest. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake Michigan to the east, Michigan to the northeast, and Lake Superior to the north. Wisconsin's capital is...
,
Saskatchewan
Saskatchewan
Saskatchewan is a prairie province in Canada, which has an area of . Saskatchewan is bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, and on the south by the U.S. states of Montana and North Dakota....
,
British Columbia
British Columbia
British Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...
,
Missouri
Missouri
Missouri is a US state located in the Midwestern United States, bordered by Iowa, Illinois, Kentucky, Tennessee, Arkansas, Oklahoma, Kansas and Nebraska. With a 2010 population of 5,988,927, Missouri is the 18th most populous state in the nation and the fifth most populous in the Midwest. It...
.
The United States Geological Survey
United States Geological Survey
The United States Geological Survey is a scientific agency of the United States government. The scientists of the USGS study the landscape of the United States, its natural resources, and the natural hazards that threaten it. The organization has four major science disciplines, concerning biology,...
reports that Common Groundsel is exotic to all 50 states and all Canadian provinces with the exception of Georgia
Georgia (U.S. state)
Georgia is a state located in the southeastern United States. It was established in 1732, the last of the original Thirteen Colonies. The state is named after King George II of Great Britain. Georgia was the fourth state to ratify the United States Constitution, on January 2, 1788...
, Kentucky
Kentucky
The Commonwealth of Kentucky is a state located in the East Central United States of America. As classified by the United States Census Bureau, Kentucky is a Southern state, more specifically in the East South Central region. Kentucky is one of four U.S. states constituted as a commonwealth...
, Massachusetts
Massachusetts
The Commonwealth of Massachusetts is a state in the New England region of the northeastern United States of America. It is bordered by Rhode Island and Connecticut to the south, New York to the west, and Vermont and New Hampshire to the north; at its east lies the Atlantic Ocean. As of the 2010...
, and Labrador
Labrador
Labrador is the distinct, northerly region of the Canadian province of Newfoundland and Labrador. It comprises the mainland portion of the province, separated from the island of Newfoundland by the Strait of Belle Isle...
.
Native
- Senecio vulgaris is considered to be native to AlgeriaAlgeriaAlgeria , officially the People's Democratic Republic of Algeria , also formally referred to as the Democratic and Popular Republic of Algeria, is a country in the Maghreb region of Northwest Africa with Algiers as its capital.In terms of land area, it is the largest country in Africa and the Arab...
, LibyaLibyaLibya is an African country in the Maghreb region of North Africa bordered by the Mediterranean Sea to the north, Egypt to the east, Sudan to the southeast, Chad and Niger to the south, and Algeria and Tunisia to the west....
, MoroccoMoroccoMorocco , officially the Kingdom of Morocco , is a country located in North Africa. It has a population of more than 32 million and an area of 710,850 km², and also primarily administers the disputed region of the Western Sahara...
, TunisiaTunisiaTunisia , officially the Tunisian RepublicThe long name of Tunisia in other languages used in the country is: , is the northernmost country in Africa. It is a Maghreb country and is bordered by Algeria to the west, Libya to the southeast, and the Mediterranean Sea to the north and east. Its area...
, GeorgiaGeorgia (country)Georgia is a sovereign state in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded to the west by the Black Sea, to the north by Russia, to the southwest by Turkey, to the south by Armenia, and to the southeast by Azerbaijan. The capital of...
, Republic of Adygea, Karachay-Cherkess RepublicKarachay-CherkessiaThe Karachay-Cherkess Republic , or Karachay-Cherkessia is a federal subject of Russia . Population: -Geography:*Area: *Borders:**internal: Krasnodar Krai , Kabardino-Balkar Republic , Stavropol Krai ....
, Kabardino-Balkaria RepublicKabardino-BalkariaThe Kabardino-Balkar Republic , or Kabardino-Balkaria , is a federal subject of Russia located in the North Caucasus. Population: -Geography:The republic is situated in the North Caucasus mountains, with plains in the northern part....
, Republic of North Ossetia-AlaniaNorth Ossetia-AlaniaThe Republic of North Ossetia–Alania is a federal subject of Russia . Its population according to the 2010 Census was 712,877.-Name:...
, Republic of IngushetiaIngushetiaThe Republic of Ingushetia is a federal subject of Russia , located in the North Caucasus region with its capital at Magas. In terms of area, the republic is the smallest of Russia's federal subjects except for the two federal cities, Moscow and Saint Petersburg...
, Chechen RepublicChechnyaThe Chechen Republic , commonly referred to as Chechnya , also spelled Chechnia or Chechenia, sometimes referred to as Ichkeria , is a federal subject of Russia . It is located in the southeastern part of Europe in the Northern Caucasus mountains. The capital of the republic is the city of Grozny...
, Republic of Dagestan, Amur OblastAmur OblastAmur Oblast is a federal subject of Russia , situated about east of Moscow on the banks of the Amur and Zeya Rivers. It shares its border with the Sakha Republic in the north, Khabarovsk Krai and the Jewish Autonomous Oblast in the east, People's Republic of China in the south, and Zabaykalsky...
, Chukotka Autonomous OkrugChukotka Autonomous OkrugChukotka Autonomous Okrug , or Chukotka , is a federal subject of Russia located in the Russian Far East.Chukotka has a population of 53,824 according to the 2002 Census, and a surface area of . The principal town and the administrative center is Anadyr...
, Jewish Autonomous OblastJewish Autonomous OblastThe Jewish Autonomous Oblast is a federal subject of Russia situated in the Russian Far East, bordering Khabarovsk Krai and Amur Oblast of Russia and Heilongjiang province of China. Its administrative center is the town of Birobidzhan....
, Kamchatka OblastKamchatka OblastKamchatka Oblast was, until being incorporated into Kamchatka Krai on July 1, 2007, a federal subject of Russia . To the north, it bordered Magadan Oblast and Chukotka Autonomous Okrug. Koryak Autonomous Okrug was located in the northern part of the oblast...
, Koryak Autonomous OkrugKoryak Autonomous OkrugKoryak Okrug , or Koryakia, is an administrative division of Kamchatka Krai, Russia. It was a federal subject of Russia from 1931 until July 1, 2007, when it merged with Kamchatka Oblast. Prior to the merger, it was called Koryak Autonomous Okrug...
, Khabarovsk KraiKhabarovsk KraiKhabarovsk Krai is a federal subject of Russia , located in the Russian Far East. It lies mostly in the basin of the lower Amur River, but also occupies a vast mountainous area along the coastline of the Sea of Okhotsk, an arm of the Pacific Ocean. The administrative center of the krai is the...
, Magadan OblastMagadan OblastMagadan Oblast is a federal subject of Russia in the Far Eastern Federal District. Its administrative center is the city of Magadan....
, Primorsky KraiPrimorsky KraiPrimorsky Krai , informally known as Primorye , is a federal subject of Russia . Primorsky means "maritime" in Russian, hence the region is sometimes referred to as Maritime Province or Maritime Territory. Its administrative center is in the city of Vladivostok...
, Sakha (Yakutia) Republic, Sakhalin OblastSakhalin OblastSakhalin Oblast is a federal subject of Russia comprising the island of Sakhalin and the Kuril Islands.The oblast has an area of 87,100 km² and a population of 546,695...
, South KoreaSouth KoreaThe Republic of Korea , , is a sovereign state in East Asia, located on the southern portion of the Korean Peninsula. It is neighbored by the People's Republic of China to the west, Japan to the east, North Korea to the north, and the East China Sea and Republic of China to the south...
, North KoreaNorth KoreaThe Democratic People’s Republic of Korea , , is a country in East Asia, occupying the northern half of the Korean Peninsula. Its capital and largest city is Pyongyang. The Korean Demilitarized Zone serves as the buffer zone between North Korea and South Korea...
, DenmarkDenmarkDenmark is a Scandinavian country in Northern Europe. The countries of Denmark and Greenland, as well as the Faroe Islands, constitute the Kingdom of Denmark . It is the southernmost of the Nordic countries, southwest of Sweden and south of Norway, and bordered to the south by Germany. Denmark...
, Faroe IslandsFaroe IslandsThe Faroe Islands are an island group situated between the Norwegian Sea and the North Atlantic Ocean, approximately halfway between Scotland and Iceland. The Faroe Islands are a self-governing territory within the Kingdom of Denmark, along with Denmark proper and Greenland...
, FinlandFinlandFinland , officially the Republic of Finland, is a Nordic country situated in the Fennoscandian region of Northern Europe. It is bordered by Sweden in the west, Norway in the north and Russia in the east, while Estonia lies to its south across the Gulf of Finland.Around 5.4 million people reside...
, IcelandIcelandIceland , described as the Republic of Iceland, is a Nordic and European island country in the North Atlantic Ocean, on the Mid-Atlantic Ridge. Iceland also refers to the main island of the country, which contains almost all the population and almost all the land area. The country has a population...
, IrelandIrelandIreland is an island to the northwest of continental Europe. It is the third-largest island in Europe and the twentieth-largest island on Earth...
, NorwayNorwayNorway , officially the Kingdom of Norway, is a Nordic unitary constitutional monarchy whose territory comprises the western portion of the Scandinavian Peninsula, Jan Mayen, and the Arctic archipelago of Svalbard and Bouvet Island. Norway has a total area of and a population of about 4.9 million...
, SwedenSwedenSweden , officially the Kingdom of Sweden , is a Nordic country on the Scandinavian Peninsula in Northern Europe. Sweden borders with Norway and Finland and is connected to Denmark by a bridge-tunnel across the Öresund....
, United KingdomUnited KingdomThe United Kingdom of Great Britain and Northern IrelandIn the United Kingdom and Dependencies, other languages have been officially recognised as legitimate autochthonous languages under the European Charter for Regional or Minority Languages...
, AustriaAustriaAustria , officially the Republic of Austria , is a landlocked country of roughly 8.4 million people in Central Europe. It is bordered by the Czech Republic and Germany to the north, Slovakia and Hungary to the east, Slovenia and Italy to the south, and Switzerland and Liechtenstein to the...
, BelgiumBelgiumBelgium , officially the Kingdom of Belgium, is a federal state in Western Europe. It is a founding member of the European Union and hosts the EU's headquarters, and those of several other major international organisations such as NATO.Belgium is also a member of, or affiliated to, many...
, Czech RepublicCzech RepublicThe Czech Republic is a landlocked country in Central Europe. The country is bordered by Poland to the northeast, Slovakia to the east, Austria to the south, and Germany to the west and northwest....
, SlovakiaSlovakiaThe Slovak Republic is a landlocked state in Central Europe. It has a population of over five million and an area of about . Slovakia is bordered by the Czech Republic and Austria to the west, Poland to the north, Ukraine to the east and Hungary to the south...
, GermanyGermanyGermany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...
, HungaryHungaryHungary , officially the Republic of Hungary , is a landlocked country in Central Europe. It is situated in the Carpathian Basin and is bordered by Slovakia to the north, Ukraine and Romania to the east, Serbia and Croatia to the south, Slovenia to the southwest and Austria to the west. The...
, NetherlandsNetherlandsThe Netherlands is a constituent country of the Kingdom of the Netherlands, located mainly in North-West Europe and with several islands in the Caribbean. Mainland Netherlands borders the North Sea to the north and west, Belgium to the south, and Germany to the east, and shares maritime borders...
, PolandPolandPoland , officially the Republic of Poland , is a country in Central Europe bordered by Germany to the west; the Czech Republic and Slovakia to the south; Ukraine, Belarus and Lithuania to the east; and the Baltic Sea and Kaliningrad Oblast, a Russian exclave, to the north...
, SwitzerlandSwitzerlandSwitzerland name of one of the Swiss cantons. ; ; ; or ), in its full name the Swiss Confederation , is a federal republic consisting of 26 cantons, with Bern as the seat of the federal authorities. The country is situated in Western Europe,Or Central Europe depending on the definition....
, BelarusBelarusBelarus , officially the Republic of Belarus, is a landlocked country in Eastern Europe, bordered clockwise by Russia to the northeast, Ukraine to the south, Poland to the west, and Lithuania and Latvia to the northwest. Its capital is Minsk; other major cities include Brest, Grodno , Gomel ,...
, EstoniaEstoniaEstonia , officially the Republic of Estonia , is a state in the Baltic region of Northern Europe. It is bordered to the north by the Gulf of Finland, to the west by the Baltic Sea, to the south by Latvia , and to the east by Lake Peipsi and the Russian Federation . Across the Baltic Sea lies...
, LatviaLatviaLatvia , officially the Republic of Latvia , is a country in the Baltic region of Northern Europe. It is bordered to the north by Estonia , to the south by Lithuania , to the east by the Russian Federation , to the southeast by Belarus and shares maritime borders to the west with Sweden...
, LithuaniaLithuaniaLithuania , officially the Republic of Lithuania is a country in Northern Europe, the biggest of the three Baltic states. It is situated along the southeastern shore of the Baltic Sea, whereby to the west lie Sweden and Denmark...
, MoldovaMoldovaMoldova , officially the Republic of Moldova is a landlocked state in Eastern Europe, located between Romania to the West and Ukraine to the North, East and South. It declared itself an independent state with the same boundaries as the preceding Moldavian Soviet Socialist Republic in 1991, as part...
, Russian Central Federal DistrictCentral Federal DistrictThe Central Federal District is one of the eight federal districts of Russia. The word "Central" is of political and historical meaning; geographically, the district is situated in the extreme west of Russia. The district spans an area of , with a population of 38,438,600 according to the 2010...
, Russian Southern Federal DistrictSouthern Federal DistrictSouthern Federal District is one of the eight federal districts of Russia. Its territory lies mostly on the Pontic-Caspian steppe. Its population was 13,856,700 according to the 2010 Census, living on an area of...
, UkraineUkraineUkraine is a country in Eastern Europe. It has an area of 603,628 km², making it the second largest contiguous country on the European continent, after Russia...
, AlbaniaAlbaniaAlbania , officially known as the Republic of Albania , is a country in Southeastern Europe, in the Balkans region. It is bordered by Montenegro to the northwest, Kosovo to the northeast, the Republic of Macedonia to the east and Greece to the south and southeast. It has a coast on the Adriatic Sea...
, BulgariaBulgariaBulgaria , officially the Republic of Bulgaria , is a parliamentary democracy within a unitary constitutional republic in Southeast Europe. The country borders Romania to the north, Serbia and Macedonia to the west, Greece and Turkey to the south, as well as the Black Sea to the east...
, GreeceGreeceGreece , officially the Hellenic Republic , and historically Hellas or the Republic of Greece in English, is a country in southeastern Europe....
, ItalyItalyItaly , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...
, RomaniaRomaniaRomania is a country located at the crossroads of Central and Southeastern Europe, on the Lower Danube, within and outside the Carpathian arch, bordering on the Black Sea...
, Bosnia and HerzegovinaBosnia and HerzegovinaBosnia and Herzegovina , sometimes called Bosnia-Herzegovina or simply Bosnia, is a country in Southern Europe, on the Balkan Peninsula. Bordered by Croatia to the north, west and south, Serbia to the east, and Montenegro to the southeast, Bosnia and Herzegovina is almost landlocked, except for the...
, CroatiaCroatiaCroatia , officially the Republic of Croatia , is a unitary democratic parliamentary republic in Europe at the crossroads of the Mitteleuropa, the Balkans, and the Mediterranean. Its capital and largest city is Zagreb. The country is divided into 20 counties and the city of Zagreb. Croatia covers ...
, the Republic of MacedoniaRepublic of MacedoniaMacedonia , officially the Republic of Macedonia , is a country located in the central Balkan peninsula in Southeast Europe. It is one of the successor states of the former Yugoslavia, from which it declared independence in 1991...
, MontenegroMontenegroMontenegro Montenegrin: Crna Gora Црна Гора , meaning "Black Mountain") is a country located in Southeastern Europe. It has a coast on the Adriatic Sea to the south-west and is bordered by Croatia to the west, Bosnia and Herzegovina to the northwest, Serbia to the northeast and Albania to the...
, SerbiaSerbiaSerbia , officially the Republic of Serbia , is a landlocked country located at the crossroads of Central and Southeast Europe, covering the southern part of the Carpathian basin and the central part of the Balkans...
, and SloveniaSloveniaSlovenia , officially the Republic of Slovenia , is a country in Central and Southeastern Europe touching the Alps and bordering the Mediterranean. Slovenia borders Italy to the west, Croatia to the south and east, Hungary to the northeast, and Austria to the north, and also has a small portion of...
, FranceFranceThe French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...
and PortugalPortugalPortugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...
.
Current
- America
- Canada: AlbertaAlbertaAlberta is a province of Canada. It had an estimated population of 3.7 million in 2010 making it the most populous of Canada's three prairie provinces...
, British ColumbiaBritish ColumbiaBritish Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...
, ManitobaManitobaManitoba is a Canadian prairie province with an area of . The province has over 110,000 lakes and has a largely continental climate because of its flat topography. Agriculture, mostly concentrated in the fertile southern and western parts of the province, is vital to the province's economy; other...
, New BrunswickNew BrunswickNew Brunswick is one of Canada's three Maritime provinces and is the only province in the federation that is constitutionally bilingual . The provincial capital is Fredericton and Saint John is the most populous city. Greater Moncton is the largest Census Metropolitan Area...
, Newfoundland and LabradorNewfoundland and LabradorNewfoundland and Labrador is the easternmost province of Canada. Situated in the country's Atlantic region, it incorporates the island of Newfoundland and mainland Labrador with a combined area of . As of April 2011, the province's estimated population is 508,400...
, Northwest TerritoriesNorthwest TerritoriesThe Northwest Territories is a federal territory of Canada.Located in northern Canada, the territory borders Canada's two other territories, Yukon to the west and Nunavut to the east, and three provinces: British Columbia to the southwest, and Alberta and Saskatchewan to the south...
and NunavutNunavutNunavut is the largest and newest federal territory of Canada; it was separated officially from the Northwest Territories on April 1, 1999, via the Nunavut Act and the Nunavut Land Claims Agreement Act, though the actual boundaries had been established in 1993...
(not collected North of the Hudson BayCanadian Arctic ArchipelagoThe Canadian Arctic Archipelago, also known as the Arctic Archipelago, is a Canadian archipelago north of the Canadian mainland in the Arctic...
), Nova ScotiaNova ScotiaNova Scotia is one of Canada's three Maritime provinces and is the most populous province in Atlantic Canada. The name of the province is Latin for "New Scotland," but "Nova Scotia" is the recognized, English-language name of the province. The provincial capital is Halifax. Nova Scotia is the...
, OntarioOntarioOntario is a province of Canada, located in east-central Canada. It is Canada's most populous province and second largest in total area. It is home to the nation's most populous city, Toronto, and the nation's capital, Ottawa....
, Prince Edward IslandPrince Edward IslandPrince Edward Island is a Canadian province consisting of an island of the same name, as well as other islands. The maritime province is the smallest in the nation in both land area and population...
, QuebecQuebecQuebec or is a province in east-central Canada. It is the only Canadian province with a predominantly French-speaking population and the only one whose sole official language is French at the provincial level....
, SaskatchewanSaskatchewanSaskatchewan is a prairie province in Canada, which has an area of . Saskatchewan is bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, and on the south by the U.S. states of Montana and North Dakota....
, YukonYukonYukon is the westernmost and smallest of Canada's three federal territories. It was named after the Yukon River. The word Yukon means "Great River" in Gwich’in....
. - Mexico: AguascalientesAguascalientesAguascalientes is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is divided in 11 municipalities and its capital city is Aguascalientes....
, Baja California NorteBaja CaliforniaBaja California officially Estado Libre y Soberano de Baja California is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is both the northernmost and westernmost state of Mexico. Before becoming a state in 1953, the area was known as the North...
, ChiapasChiapasChiapas officially Estado Libre y Soberano de Chiapas is one of the 31 states that, with the Federal District, comprise the 32 Federal Entities of Mexico. It is divided in 118 municipalities and its capital city is Tuxtla Gutierrez. Other important cites in Chiapas include San Cristóbal de las...
, CoahuilaCoahuilaCoahuila, formally Coahuila de Zaragoza , officially Estado Libre y Soberano de Coahuila de Zaragoza is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico...
, Distrito FederalMexico CityMexico City is the Federal District , capital of Mexico and seat of the federal powers of the Mexican Union. It is a federal entity within Mexico which is not part of any one of the 31 Mexican states but belongs to the federation as a whole...
, Hidalgo, JaliscoJaliscoJalisco officially Estado Libre y Soberano de Jalisco is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is located in Western Mexico and divided in 125 municipalities and its capital city is Guadalajara.It is one of the more important states...
, Estado de México, Nuevo LeónNuevo LeónNuevo León It is located in Northeastern Mexico. It is bordered by the states of Tamaulipas to the north and east, San Luis Potosí to the south, and Coahuila to the west. To the north, Nuevo León has a 15 kilometer stretch of the U.S.-Mexico border adjacent to the U.S...
, PueblaPueblaPuebla officially Estado Libre y Soberano de Puebla is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is divided in 217 municipalities and its capital city is Puebla....
, TlaxcalaTlaxcalaTlaxcala officially Estado Libre y Soberano de Tlaxcala is one of the 31 states which along with the Federal District comprise the 32 federative entities of Mexico. It is divided into 60 municipalities and its capital city is Tlaxcala....
, VeracruzVeracruzVeracruz, formally Veracruz de Ignacio de la Llave officially Estado Libre y Soberano de Veracruz de Ignacio de la Llave , is one of the 31 states that, along with the Federal District, comprise the 32 federative entities of Mexico. It is divided in 212 municipalities and its capital city is... - North America: United States of AmericaUnited StatesThe United States of America is a federal constitutional republic comprising fifty states and a federal district...
, GreenlandGreenlandGreenland is an autonomous country within the Kingdom of Denmark, located between the Arctic and Atlantic Oceans, east of the Canadian Arctic Archipelago. Though physiographically a part of the continent of North America, Greenland has been politically and culturally associated with Europe for...
, Saint Pierre and Miquelon - South America: ArgentinaArgentinaArgentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...
, BoliviaBoliviaBolivia officially known as Plurinational State of Bolivia , is a landlocked country in central South America. It is the poorest country in South America...
, ChileChileChile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...
, EcuadorEcuadorEcuador , officially the Republic of Ecuador is a representative democratic republic in South America, bordered by Colombia on the north, Peru on the east and south, and by the Pacific Ocean to the west. It is one of only two countries in South America, along with Chile, that do not have a border...
, PeruPeruPeru , officially the Republic of Peru , is a country in western South America. It is bordered on the north by Ecuador and Colombia, on the east by Brazil, on the southeast by Bolivia, on the south by Chile, and on the west by the Pacific Ocean....
- Canada: Alberta
- Africa
- Northern Africa: AlgeriaAlgeriaAlgeria , officially the People's Democratic Republic of Algeria , also formally referred to as the Democratic and Popular Republic of Algeria, is a country in the Maghreb region of Northwest Africa with Algiers as its capital.In terms of land area, it is the largest country in Africa and the Arab...
, EgyptEgyptEgypt , officially the Arab Republic of Egypt, Arabic: , is a country mainly in North Africa, with the Sinai Peninsula forming a land bridge in Southwest Asia. Egypt is thus a transcontinental country, and a major power in Africa, the Mediterranean Basin, the Middle East and the Muslim world...
including Sinai, LibyaLibyaLibya is an African country in the Maghreb region of North Africa bordered by the Mediterranean Sea to the north, Egypt to the east, Sudan to the southeast, Chad and Niger to the south, and Algeria and Tunisia to the west....
, MoroccoMoroccoMorocco , officially the Kingdom of Morocco , is a country located in North Africa. It has a population of more than 32 million and an area of 710,850 km², and also primarily administers the disputed region of the Western Sahara...
and Spanish MoroccoSpanish MoroccoThe Spanish protectorate of Morocco was the area of Morocco under colonial rule by the Spanish Empire, established by the Treaty of Fez in 1912 and ending in 1956, when both France and Spain recognized Moroccan independence.-Territorial borders:...
, TunisiaTunisiaTunisia , officially the Tunisian RepublicThe long name of Tunisia in other languages used in the country is: , is the northernmost country in Africa. It is a Maghreb country and is bordered by Algeria to the west, Libya to the southeast, and the Mediterranean Sea to the north and east. Its area...
- Northern Africa: Algeria
- Asia
- Western Asia: IranIranIran , officially the Islamic Republic of Iran , is a country in Southern and Western Asia. The name "Iran" has been in use natively since the Sassanian era and came into use internationally in 1935, before which the country was known to the Western world as Persia...
, IsraelIsraelThe State of Israel is a parliamentary republic located in the Middle East, along the eastern shore of the Mediterranean Sea...
with the Palestinian territoriesPalestinian territoriesThe Palestinian territories comprise the West Bank and the Gaza Strip. Since the Palestinian Declaration of Independence in 1988, the region is today recognized by three-quarters of the world's countries as the State of Palestine or simply Palestine, although this status is not recognized by the...
, LebanonLebanonLebanon , officially the Republic of LebanonRepublic of Lebanon is the most common term used by Lebanese government agencies. The term Lebanese Republic, a literal translation of the official Arabic and French names that is not used in today's world. Arabic is the most common language spoken among...
and SyriaSyriaSyria , officially the Syrian Arab Republic , is a country in Western Asia, bordering Lebanon and the Mediterranean Sea to the West, Turkey to the north, Iraq to the east, Jordan to the south, and Israel to the southwest....
. - Caucasus: Adygea, AzerbaijanAzerbaijanAzerbaijan , officially the Republic of Azerbaijan is the largest country in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded by the Caspian Sea to the east, Russia to the north, Georgia to the northwest, Armenia to the west, and Iran to...
, ChechnyaChechnyaThe Chechen Republic , commonly referred to as Chechnya , also spelled Chechnia or Chechenia, sometimes referred to as Ichkeria , is a federal subject of Russia . It is located in the southeastern part of Europe in the Northern Caucasus mountains. The capital of the republic is the city of Grozny...
, Dagestan, GeorgiaGeorgia (country)Georgia is a sovereign state in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded to the west by the Black Sea, to the north by Russia, to the southwest by Turkey, to the south by Armenia, and to the southeast by Azerbaijan. The capital of...
, IngushetiaIngushetiaThe Republic of Ingushetia is a federal subject of Russia , located in the North Caucasus region with its capital at Magas. In terms of area, the republic is the smallest of Russia's federal subjects except for the two federal cities, Moscow and Saint Petersburg...
, Kabardino-BalkariaKabardino-BalkariaThe Kabardino-Balkar Republic , or Kabardino-Balkaria , is a federal subject of Russia located in the North Caucasus. Population: -Geography:The republic is situated in the North Caucasus mountains, with plains in the northern part....
, KalmykiaKalmykiaThe Republic of Kalmykia is a federal subject of Russia . Population: It is the only Buddhist region in Europe. It has also become well-known as an international chess mecca because its former President, Kirsan Ilyumzhinov, is the head of the International Chess Federation .-Geography:*Area:...
, Karachay-CherkessiaKarachay-CherkessiaThe Karachay-Cherkess Republic , or Karachay-Cherkessia is a federal subject of Russia . Population: -Geography:*Area: *Borders:**internal: Krasnodar Krai , Kabardino-Balkar Republic , Stavropol Krai ....
, North Ossetia-AlaniaNorth Ossetia-AlaniaThe Republic of North Ossetia–Alania is a federal subject of Russia . Its population according to the 2010 Census was 712,877.-Name:... - Northwestern Asia: Arkhangelsk OblastArkhangelsk OblastArkhangelsk Oblast is a federal subject of Russia . It includes the Arctic archipelagos of Franz Josef Land and Novaya Zemlya, as well as the Solovetsky Islands in the White Sea....
including Nenets Autonomous OkrugNenets Autonomous OkrugNenets Autonomous Okrug is a federal subject of Russia .It has an area of 176,700 km2 and population of 42,628 as of the preliminary results of the 2010 Census , 21,296 of whom live in Naryan-Mar, the administrative center.-Geography and ecology:The arctic ecology of this...
and Novaya ZemlyaNovaya ZemlyaNovaya Zemlya , also known in Dutch as Nova Zembla and in Norwegian as , is an archipelago in the Arctic Ocean in the north of Russia and the extreme northeast of Europe, the easternmost point of Europe lying at Cape Flissingsky on the northern island...
, Astrakhan OblastAstrakhan OblastAstrakhan Oblast is a federal subject of Russia . Its administrative center is the city of Astrakhan.-Demographics:Population: Ethnic groups...
, BashkortostanBashkortostanThe Republic of Bashkortostan , also known as Bashkiria is a federal subject of Russia . It is located between the Volga River and the Ural Mountains. Its capital is the city of Ufa...
, Belgorod OblastBelgorod OblastBelgorod Oblast is a federal subjects of Russia . Its administrative center is the city of Belgorod. Population: 1,532,670 .-History:...
, Bryansk OblastBryansk OblastBryansk Oblast is a federal subject of Russia . Its administrative center is the city of Bryansk. Population: 1,278,087 .-History:...
, Chuvashia, Lipetsk OblastLipetsk OblastLipetsk Oblast is a federal subject of Russia which was formed on January 6, 1954. Its administrative center is the city of Lipetsk...
, Kaliningrad OblastKaliningrad OblastKaliningrad Oblast is a federal subject of Russia situated on the Baltic coast. It has a population of The oblast forms the westernmost part of the Russian Federation, but it has no land connection to the rest of Russia. Since its creation it has been an exclave of the Russian SFSR and then the...
, Kaluga OblastKaluga OblastKaluga Oblast is a federal subject of Russia . Its administrative center is the city of Kaluga.-Geography:Kaluga Oblast is located in the central part of the East European Plain. The Smolensk Highland lays in the western and north-western part of the oblast, while the Central Russian Highland -...
, Novgorod OblastNovgorod OblastNovgorod Oblast is a federal subject of Russia , located between Moscow and Saint Petersburg. Its administrative center is the city of Veliky Novgorod. Some of the oldest Russian cities, including Veliky Novgorod and Staraya Russa, are located there...
, Republic of KareliaRepublic of KareliaThe Republic of Karelia is a federal subject of Russia .-Geography:The republic is located in the northwestern part of Russia, taking intervening position between the basins of White and Baltic seas...
, Kirov OblastKirov OblastKirov Oblast is a federal subject of Russia . Its administrative center is the city of Kirov. Population: -History:In the late 19th and early 20th centuries, Vyatka remained a place of exile for opponents of the tsarist regime, including many prominent revolutionary figures.In 1920, a number of...
, Komi RepublicKomi RepublicThe Komi Republic is a federal subject of Russia .-Geography:The republic is situated to the west of the Ural mountains, in the north-east of the East European Plain...
, Kursk OblastKursk OblastKursk Oblast is a federal subject of Russia . Its administrative center is the city of Kursk.-Geography:The oblast occupies the southern slopes of the middle-Russian plateau, and its average elevation is from 177 to 225 meters . The surface is hilly, and intersected by ravines...
, MordoviaMordoviaThe Republic of Mordovia , also known as Mordvinia, is a federal subject of Russia . Its capital is the city of Saransk. Population: -Geography:The republic is located in the eastern part of the East European Plain of Russia...
, Murmansk OblastMurmansk OblastMurmansk Oblast is a federal subject of Russia , located in the northwestern part of Russia. Its administrative center is the city of Murmansk.-Geography:...
, Orenburg OblastOrenburg OblastOrenburg Oblast is a federal subject of Russia . Its administrative center is the city of Orenburg. From 1938 to 1957, it bore the name Chkalov Oblast in honor of Valery Chkalov...
, Penza OblastPenza Oblast-External links:* *...
, Perm KraiPerm KraiPerm Krai is a federal subject of Russia that came into existence on December 1, 2005 as a result of the 2004 referendum on the merger of Perm Oblast and Komi-Permyak Autonomous Okrug. The city of Perm became the administrative center of the new federal subject...
, Pskov OblastPskov OblastPskov Oblast is a federal subject of Russia . Pskov Oblast borders the countries of Estonia and Latvia, as well as Belarus. It is the westernmost federal subject of contiguous Russia . Its major cities are the administrative center Pskov and Velikiye Luki . Area: 55,300 km²...
, Rostov OblastRostov OblastRostov Oblast is a federal subject of Russia , located in the Southern Federal District. Rostov Oblast has an area of and a population of making it the sixth most populous federal subject in Russia...
, Ryazan OblastRyazan OblastRyazan Oblast is a federal subject of Russia . Its administrative center is the city of Ryazan, which is the oblast's largest city. Population: -Geography:...
, Saint PetersburgSaint PetersburgSaint Petersburg is a city and a federal subject of Russia located on the Neva River at the head of the Gulf of Finland on the Baltic Sea...
, Samara OblastSamara OblastSamara Oblast is a federal subject of Russia . Its administrative center is the city of Samara. Population: In 1936–1990, it was known as Kuybyshev Oblast , after the Soviet name of Samara .-Demographics:Population:...
, Saratov OblastSaratov OblastSaratov Oblast is a federal subject of Russia , located in the Volga Federal District. Its administrative center is the city of Saratov. Population: -Demographics:Population:...
, Tambov OblastTambov OblastTambov Oblast is a federal subject of Russia . Its administrative center is the city of Tambov. Population: down from 1,178,443 recorded by the 2002 Census.Tambov Oblast is situated in forest steppe.-Birth rate:...
, TatarstanTatarstanThe Republic of Tatarstan is a federal subject of Russia located in the Volga Federal District. Its capital is the city of Kazan, which is one of Russia's largest and most prosperous cities. The republic borders with Kirov, Ulyanovsk, Samara, and Orenburg Oblasts, and with the Mari El, Udmurt,...
, Tula OblastTula OblastTula Oblast is a federal subject of Russia with its present borders formed on September 26, 1937. Its administrative center is the city of Tula. The oblast has an area of and a population of 1,553,874...
, UdmurtiaUdmurtiaThe Udmurt Republic , or Udmurtia is a federal subject of Russia . Its capital is the city of Izhevsk. Population: -History:...
, Volgograd OblastVolgograd OblastVolgograd Oblast is a federal subject of Russia . Its administrative center is the city of Volgograd. Population: -Geography:*Area: 113,900 km²;*Borders length: 2221,9 km²....
, Vologda OblastVologda OblastVologda Oblast is a federal subject of Russia . Its administrative center is Vologda. The largest city is Cherepovets.Vologda Oblast is rich in historic monuments, such as the magnificent Kirillo-Belozersky Monastery, Ferapontov Convent , medieval towns of Velikiy Ustyug and Belozersk, baroque...
, Voronezh OblastVoronezh OblastVoronezh Oblast is a federal subject of Russia . It was established on June 13, 1934.-Main rivers:*Don*Voronezh*Bityug*Khopyor-Economy:...
, - Siberia: Altai KraiAltai KraiAltai Krai is a federal subject of Russia . It borders with, clockwise from the south, Kazakhstan, Novosibirsk and Kemerovo Oblasts, and the Altai Republic. The krai's administrative center is the city of Barnaul...
, Buryatia, Chelyabinsk OblastChelyabinsk Oblast-External links:*...
, Chita OblastChita OblastChita Oblast was a federal subject of Russia in southeast Siberia, Russia. Its administrative center was the city of Chita. It had extensive international borders with China and Mongolia and internal borders with Irkutsk and Amur Oblasts, as well as with the Buryat and the Sakha Republics. Its...
, Irkutsk OblastIrkutsk OblastIrkutsk Oblast is a federal subject of Russia , located in southeastern Siberia in the basins of Angara River, Lena, and Nizhnyaya Tunguska Rivers. The administrative center is the city of Irkutsk. Population: -History:...
, Kemerovo OblastKemerovo OblastKemerovo Oblast , also known as Kuzbass after the Kuznetsk Basin, is a federal subject of Russia , located in southwestern Siberia, where the West Siberian Plain meets the South Siberian mountains...
, Krasnoyarsk KraiKrasnoyarsk KraiKrasnoyarsk Krai is a federal subject of Russia . It is the second largest federal subject after the Sakha Republic, and Russia's largest krai, occupying an area of , which is 13% of the country's total territory. The administrative center of the krai is the city of Krasnoyarsk...
, parts of Kurgan OblastKurgan OblastKurgan Oblast is a federal subject of Russia . Its administrative center is the city of Kurgan. Population: -History:The oblast was formed on February 6, 1943, just when the Soviet Army decisively defeated Hitler's forces near Stalingrad...
, Novosibirsk OblastNovosibirsk OblastNovosibirsk Oblast is a federal subject of Russia located in southwestern Siberia. Its administrative and economic center is the city of Novosibirsk. Population: -Overview:...
, Omsk OblastOmsk OblastOmsk Oblast is a federal subject of Russia , located in southwestern Siberia. The oblast has an area of and a population of with the majority, 1.15 million, living in Omsk, the administrative center....
, Sverdlovsk OblastSverdlovsk OblastSverdlovsk Oblast is a federal subject of Russia located in the Urals Federal District. Its administrative center is the city of Yekaterinburg formerly known as Sverdlovsk. Population: -Geography:...
, Tomsk OblastTomsk OblastTomsk Oblast is a federal subject of Russia . It lies in the southeastern West Siberian Plain, in the southwest of the Siberian Federal District. Its administrative center is the city of Tomsk. Population:...
, TuvaTuvaThe Tyva Republic , or Tuva , is a federal subject of Russia . It lies in the geographical center of Asia, in southern Siberia. The republic borders with the Altai Republic, the Republic of Khakassia, Krasnoyarsk Krai, Irkutsk Oblast, and the Republic of Buryatia in Russia and with Mongolia to the...
, Tyumen OblastTyumen OblastTyumen Oblast is a federal subject of Russia . Its administrative center is the city of Tyumen. The oblast has administrative jurisdiction over two autonomous okrugs—Khanty-Mansi Autonomous Okrug and Yamalo-Nenets Autonomous Okrug. Tyumen is the largest city, with over half a million inhabitants...
, Ulyanovsk OblastUlyanovsk OblastUlyanovsk Oblast is a federal subject of Russia . It is located in the Volga Federal District. Its administrative center is the city of Ulyanovsk...
, Yamal-Nenets, - Soviet Far East: Amur OblastAmur OblastAmur Oblast is a federal subject of Russia , situated about east of Moscow on the banks of the Amur and Zeya Rivers. It shares its border with the Sakha Republic in the north, Khabarovsk Krai and the Jewish Autonomous Oblast in the east, People's Republic of China in the south, and Zabaykalsky...
, Chukotka Autonomous OkrugChukotka Autonomous OkrugChukotka Autonomous Okrug , or Chukotka , is a federal subject of Russia located in the Russian Far East.Chukotka has a population of 53,824 according to the 2002 Census, and a surface area of . The principal town and the administrative center is Anadyr...
, Jewish Autonomous OblastJewish Autonomous OblastThe Jewish Autonomous Oblast is a federal subject of Russia situated in the Russian Far East, bordering Khabarovsk Krai and Amur Oblast of Russia and Heilongjiang province of China. Its administrative center is the town of Birobidzhan....
, Kamchatka OblastKamchatka OblastKamchatka Oblast was, until being incorporated into Kamchatka Krai on July 1, 2007, a federal subject of Russia . To the north, it bordered Magadan Oblast and Chukotka Autonomous Okrug. Koryak Autonomous Okrug was located in the northern part of the oblast...
, Koryak Autonomous OkrugKoryak Autonomous OkrugKoryak Okrug , or Koryakia, is an administrative division of Kamchatka Krai, Russia. It was a federal subject of Russia from 1931 until July 1, 2007, when it merged with Kamchatka Oblast. Prior to the merger, it was called Koryak Autonomous Okrug...
, Khabarovsk KraiKhabarovsk KraiKhabarovsk Krai is a federal subject of Russia , located in the Russian Far East. It lies mostly in the basin of the lower Amur River, but also occupies a vast mountainous area along the coastline of the Sea of Okhotsk, an arm of the Pacific Ocean. The administrative center of the krai is the...
, Magadan OblastMagadan OblastMagadan Oblast is a federal subject of Russia in the Far Eastern Federal District. Its administrative center is the city of Magadan....
, Primorsky KraiPrimorsky KraiPrimorsky Krai , informally known as Primorye , is a federal subject of Russia . Primorsky means "maritime" in Russian, hence the region is sometimes referred to as Maritime Province or Maritime Territory. Its administrative center is in the city of Vladivostok...
, Sakha (Yakutia) Republic, Sakhalin OblastSakhalin OblastSakhalin Oblast is a federal subject of Russia comprising the island of Sakhalin and the Kuril Islands.The oblast has an area of 87,100 km² and a population of 546,695... - China: Inner MongoliaInner MongoliaInner Mongolia is an autonomous region of the People's Republic of China, located in the northern region of the country. Inner Mongolia shares an international border with the countries of Mongolia and the Russian Federation...
, HeilongjiangHeilongjiangFor the river known in Mandarin as Heilong Jiang, see Amur River' is a province of the People's Republic of China located in the northeastern part of the country. "Heilongjiang" literally means Black Dragon River, which is the Chinese name for the Amur. The one-character abbreviation is 黑...
, JilinJilinJilin , is a province of the People's Republic of China located in the northeastern part of the country. Jilin borders North Korea and Russia to the east, Heilongjiang to the north, Liaoning to the south, and Inner Mongolia to the west...
, LiaoningLiaoning' is a province of the People's Republic of China, located in the northeast of the country. Its one-character abbreviation is "辽" , a name taken from the Liao River that flows through the province. "Níng" means "peace"...
, HebeiHebei' is a province of the People's Republic of China in the North China region. Its one-character abbreviation is "" , named after Ji Province, a Han Dynasty province that included what is now southern Hebei...
and BeijingBeijingBeijing , also known as Peking , is the capital of the People's Republic of China and one of the most populous cities in the world, with a population of 19,612,368 as of 2010. The city is the country's political, cultural, and educational center, and home to the headquarters for most of China's... - Eastern Asia: South KoreaSouth KoreaThe Republic of Korea , , is a sovereign state in East Asia, located on the southern portion of the Korean Peninsula. It is neighbored by the People's Republic of China to the west, Japan to the east, North Korea to the north, and the East China Sea and Republic of China to the south...
, North KoreaNorth KoreaThe Democratic People’s Republic of Korea , , is a country in East Asia, occupying the northern half of the Korean Peninsula. Its capital and largest city is Pyongyang. The Korean Demilitarized Zone serves as the buffer zone between North Korea and South Korea...
, HokkaidōHokkaido, formerly known as Ezo, Yezo, Yeso, or Yesso, is Japan's second largest island; it is also the largest and northernmost of Japan's 47 prefectural-level subdivisions. The Tsugaru Strait separates Hokkaido from Honshu, although the two islands are connected by the underwater railway Seikan Tunnel...
, HonshūHonshuis the largest island of Japan. The nation's main island, it is south of Hokkaido across the Tsugaru Strait, north of Shikoku across the Inland Sea, and northeast of Kyushu across the Kanmon Strait...
, ShikokuShikokuis the smallest and least populous of the four main islands of Japan, located south of Honshū and east of the island of Kyūshū. Its ancient names include Iyo-no-futana-shima , Iyo-shima , and Futana-shima...
, KyūshūKyushuis the third largest island of Japan and most southwesterly of its four main islands. Its alternate ancient names include , , and . The historical regional name is referred to Kyushu and its surrounding islands....
, Okinawa IslandOkinawa IslandOkinawa Island is the largest of the Okinawa Islands and the Ryukyu Islands of Japan, and is home to Naha, the capital of Okinawa Prefecture. The island has an area of...
- Western Asia: Iran
- Europe
- Northern Europe: Channel IslandsChannel IslandsThe Channel Islands are an archipelago of British Crown Dependencies in the English Channel, off the French coast of Normandy. They include two separate bailiwicks: the Bailiwick of Guernsey and the Bailiwick of Jersey...
, DenmarkDenmarkDenmark is a Scandinavian country in Northern Europe. The countries of Denmark and Greenland, as well as the Faroe Islands, constitute the Kingdom of Denmark . It is the southernmost of the Nordic countries, southwest of Sweden and south of Norway, and bordered to the south by Germany. Denmark...
, EstoniaEstoniaEstonia , officially the Republic of Estonia , is a state in the Baltic region of Northern Europe. It is bordered to the north by the Gulf of Finland, to the west by the Baltic Sea, to the south by Latvia , and to the east by Lake Peipsi and the Russian Federation . Across the Baltic Sea lies...
, Faeroe Islands, FinlandFinlandFinland , officially the Republic of Finland, is a Nordic country situated in the Fennoscandian region of Northern Europe. It is bordered by Sweden in the west, Norway in the north and Russia in the east, while Estonia lies to its south across the Gulf of Finland.Around 5.4 million people reside...
, IcelandIcelandIceland , described as the Republic of Iceland, is a Nordic and European island country in the North Atlantic Ocean, on the Mid-Atlantic Ridge. Iceland also refers to the main island of the country, which contains almost all the population and almost all the land area. The country has a population...
, IrelandIrelandIreland is an island to the northwest of continental Europe. It is the third-largest island in Europe and the twentieth-largest island on Earth...
with Northern IrelandNorthern IrelandNorthern Ireland is one of the four countries of the United Kingdom. Situated in the north-east of the island of Ireland, it shares a border with the Republic of Ireland to the south and west...
, NorwayNorwayNorway , officially the Kingdom of Norway, is a Nordic unitary constitutional monarchy whose territory comprises the western portion of the Scandinavian Peninsula, Jan Mayen, and the Arctic archipelago of Svalbard and Bouvet Island. Norway has a total area of and a population of about 4.9 million...
, SwedenSwedenSweden , officially the Kingdom of Sweden , is a Nordic country on the Scandinavian Peninsula in Northern Europe. Sweden borders with Norway and Finland and is connected to Denmark by a bridge-tunnel across the Öresund....
, United KingdomUnited KingdomThe United Kingdom of Great Britain and Northern IrelandIn the United Kingdom and Dependencies, other languages have been officially recognised as legitimate autochthonous languages under the European Charter for Regional or Minority Languages... - Middle Europe: AustriaAustriaAustria , officially the Republic of Austria , is a landlocked country of roughly 8.4 million people in Central Europe. It is bordered by the Czech Republic and Germany to the north, Slovakia and Hungary to the east, Slovenia and Italy to the south, and Switzerland and Liechtenstein to the...
, Czech RepublicCzech RepublicThe Czech Republic is a landlocked country in Central Europe. The country is bordered by Poland to the northeast, Slovakia to the east, Austria to the south, and Germany to the west and northwest....
, GermanyGermanyGermany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...
, PolandPolandPoland , officially the Republic of Poland , is a country in Central Europe bordered by Germany to the west; the Czech Republic and Slovakia to the south; Ukraine, Belarus and Lithuania to the east; and the Baltic Sea and Kaliningrad Oblast, a Russian exclave, to the north...
, SlovakiaSlovakiaThe Slovak Republic is a landlocked state in Central Europe. It has a population of over five million and an area of about . Slovakia is bordered by the Czech Republic and Austria to the west, Poland to the north, Ukraine to the east and Hungary to the south...
, SloveniaSloveniaSlovenia , officially the Republic of Slovenia , is a country in Central and Southeastern Europe touching the Alps and bordering the Mediterranean. Slovenia borders Italy to the west, Croatia to the south and east, Hungary to the northeast, and Austria to the north, and also has a small portion of...
, - East Europe: BelarusBelarusBelarus , officially the Republic of Belarus, is a landlocked country in Eastern Europe, bordered clockwise by Russia to the northeast, Ukraine to the south, Poland to the west, and Lithuania and Latvia to the northwest. Its capital is Minsk; other major cities include Brest, Grodno , Gomel ,...
, HungaryHungaryHungary , officially the Republic of Hungary , is a landlocked country in Central Europe. It is situated in the Carpathian Basin and is bordered by Slovakia to the north, Ukraine and Romania to the east, Serbia and Croatia to the south, Slovenia to the southwest and Austria to the west. The...
, LatviaLatviaLatvia , officially the Republic of Latvia , is a country in the Baltic region of Northern Europe. It is bordered to the north by Estonia , to the south by Lithuania , to the east by the Russian Federation , to the southeast by Belarus and shares maritime borders to the west with Sweden...
, LithuaniaLithuaniaLithuania , officially the Republic of Lithuania is a country in Northern Europe, the biggest of the three Baltic states. It is situated along the southeastern shore of the Baltic Sea, whereby to the west lie Sweden and Denmark...
, MoldaviaMoldaviaMoldavia is a geographic and historical region and former principality in Eastern Europe, corresponding to the territory between the Eastern Carpathians and the Dniester river...
, UkraineUkraineUkraine is a country in Eastern Europe. It has an area of 603,628 km², making it the second largest contiguous country on the European continent, after Russia... - Southeastern Europe: Bosnia and HerzegovinaBosnia and HerzegovinaBosnia and Herzegovina , sometimes called Bosnia-Herzegovina or simply Bosnia, is a country in Southern Europe, on the Balkan Peninsula. Bordered by Croatia to the north, west and south, Serbia to the east, and Montenegro to the southeast, Bosnia and Herzegovina is almost landlocked, except for the...
, BulgariaBulgariaBulgaria , officially the Republic of Bulgaria , is a parliamentary democracy within a unitary constitutional republic in Southeast Europe. The country borders Romania to the north, Serbia and Macedonia to the west, Greece and Turkey to the south, as well as the Black Sea to the east...
, CroatiaCroatiaCroatia , officially the Republic of Croatia , is a unitary democratic parliamentary republic in Europe at the crossroads of the Mitteleuropa, the Balkans, and the Mediterranean. Its capital and largest city is Zagreb. The country is divided into 20 counties and the city of Zagreb. Croatia covers ...
, Republic of MacedoniaRepublic of MacedoniaMacedonia , officially the Republic of Macedonia , is a country located in the central Balkan peninsula in Southeast Europe. It is one of the successor states of the former Yugoslavia, from which it declared independence in 1991...
, MontenegroMontenegroMontenegro Montenegrin: Crna Gora Црна Гора , meaning "Black Mountain") is a country located in Southeastern Europe. It has a coast on the Adriatic Sea to the south-west and is bordered by Croatia to the west, Bosnia and Herzegovina to the northwest, Serbia to the northeast and Albania to the...
, RomaniaRomaniaRomania is a country located at the crossroads of Central and Southeastern Europe, on the Lower Danube, within and outside the Carpathian arch, bordering on the Black Sea...
, SerbiaSerbiaSerbia , officially the Republic of Serbia , is a landlocked country located at the crossroads of Central and Southeast Europe, covering the southern part of the Carpathian basin and the central part of the Balkans... - Southwestern Europe: AndorraAndorraAndorra , officially the Principality of Andorra , also called the Principality of the Valleys of Andorra, , is a small landlocked country in southwestern Europe, located in the eastern Pyrenees mountains and bordered by Spain and France. It is the sixth smallest nation in Europe having an area of...
, AzoresAzoresThe Archipelago of the Azores is composed of nine volcanic islands situated in the middle of the North Atlantic Ocean, and is located about west from Lisbon and about east from the east coast of North America. The islands, and their economic exclusion zone, form the Autonomous Region of the...
, Balearic IslandsBalearic IslandsThe Balearic Islands are an archipelago of Spain in the western Mediterranean Sea, near the eastern coast of the Iberian Peninsula.The four largest islands are: Majorca, Minorca, Ibiza and Formentera. The archipelago forms an autonomous community and a province of Spain with Palma as the capital...
, Canary IslandsCanary IslandsThe Canary Islands , also known as the Canaries , is a Spanish archipelago located just off the northwest coast of mainland Africa, 100 km west of the border between Morocco and the Western Sahara. The Canaries are a Spanish autonomous community and an outermost region of the European Union...
, GibraltarGibraltarGibraltar is a British overseas territory located on the southern end of the Iberian Peninsula at the entrance of the Mediterranean. A peninsula with an area of , it has a northern border with Andalusia, Spain. The Rock of Gibraltar is the major landmark of the region...
, PortugalPortugalPortugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...
, SpainSpainSpain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula... - West Europe: BelgiumBelgiumBelgium , officially the Kingdom of Belgium, is a federal state in Western Europe. It is a founding member of the European Union and hosts the EU's headquarters, and those of several other major international organisations such as NATO.Belgium is also a member of, or affiliated to, many...
, FranceFranceThe French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...
, LiechtensteinLiechtensteinThe Principality of Liechtenstein is a doubly landlocked alpine country in Central Europe, bordered by Switzerland to the west and south and by Austria to the east. Its area is just over , and it has an estimated population of 35,000. Its capital is Vaduz. The biggest town is Schaan...
, LuxembourgLuxembourgLuxembourg , officially the Grand Duchy of Luxembourg , is a landlocked country in western Europe, bordered by Belgium, France, and Germany. It has two principal regions: the Oesling in the North as part of the Ardennes massif, and the Gutland in the south...
, NetherlandsNetherlandsThe Netherlands is a constituent country of the Kingdom of the Netherlands, located mainly in North-West Europe and with several islands in the Caribbean. Mainland Netherlands borders the North Sea to the north and west, Belgium to the south, and Germany to the east, and shares maritime borders...
, SwitzerlandSwitzerlandSwitzerland name of one of the Swiss cantons. ; ; ; or ), in its full name the Swiss Confederation , is a federal republic consisting of 26 cantons, with Bern as the seat of the federal authorities. The country is situated in Western Europe,Or Central Europe depending on the definition....
, - South Europe: AlbaniaAlbaniaAlbania , officially known as the Republic of Albania , is a country in Southeastern Europe, in the Balkans region. It is bordered by Montenegro to the northwest, Kosovo to the northeast, the Republic of Macedonia to the east and Greece to the south and southeast. It has a coast on the Adriatic Sea...
, ChiosChiosChios is the fifth largest of the Greek islands, situated in the Aegean Sea, seven kilometres off the Asia Minor coast. The island is separated from Turkey by the Chios Strait. The island is noted for its strong merchant shipping community, its unique mastic gum and its medieval villages...
, CorsicaCorsicaCorsica is an island in the Mediterranean Sea. It is located west of Italy, southeast of the French mainland, and north of the island of Sardinia....
, CreteCreteCrete is the largest and most populous of the Greek islands, the fifth largest island in the Mediterranean Sea, and one of the thirteen administrative regions of Greece. It forms a significant part of the economy and cultural heritage of Greece while retaining its own local cultural traits...
, CyprusCyprusCyprus , officially the Republic of Cyprus , is a Eurasian island country, member of the European Union, in the Eastern Mediterranean, east of Greece, south of Turkey, west of Syria and north of Egypt. It is the third largest island in the Mediterranean Sea.The earliest known human activity on the...
, DodecaneseDodecaneseThe Dodecanese are a group of 12 larger plus 150 smaller Greek islands in the Aegean Sea, of which 26 are inhabited. Τhis island group generally defines the eastern limit of the Sea of Crete. They belong to the Southern Sporades island group...
, GreeceGreeceGreece , officially the Hellenic Republic , and historically Hellas or the Republic of Greece in English, is a country in southeastern Europe....
, Ikaria, ItalyItalyItaly , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...
, KarpathosKarpathosKarpathos is the second largest of the Greek Dodecanese islands, in the southeastern Aegean Sea. Together with the neighboring smaller Saria Island it forms the municipality Karpathos, which is part of the Karpathos peripheral unit. From its remote position Karpathos has preserved many...
, Lesbos, Samos IslandSamos IslandSamos is a Greek island in the eastern Aegean Sea, south of Chios, north of Patmos and the Dodecanese, and off the coast of Asia Minor, from which it is separated by the -wide Mycale Strait. It is also a separate regional unit of the North Aegean region, and the only municipality of the regional...
, San MarinoSan MarinoSan Marino, officially the Republic of San Marino , is a state situated on the Italian Peninsula on the eastern side of the Apennine Mountains. It is an enclave surrounded by Italy. Its size is just over with an estimated population of over 30,000. Its capital is the City of San Marino...
, SardiniaSardiniaSardinia is the second-largest island in the Mediterranean Sea . It is an autonomous region of Italy, and the nearest land masses are the French island of Corsica, the Italian Peninsula, Sicily, Tunisia and the Spanish Balearic Islands.The name Sardinia is from the pre-Roman noun *sard[],...
, SicilySicilySicily is a region of Italy, and is the largest island in the Mediterranean Sea. Along with the surrounding minor islands, it constitutes an autonomous region of Italy, the Regione Autonoma Siciliana Sicily has a rich and unique culture, especially with regard to the arts, music, literature,...
with MaltaMaltaMalta , officially known as the Republic of Malta , is a Southern European country consisting of an archipelago situated in the centre of the Mediterranean, south of Sicily, east of Tunisia and north of Libya, with Gibraltar to the west and Alexandria to the east.Malta covers just over in...
, Asiatic Turkey and Turkey-in-Europe, Vatican CityVatican CityVatican City , or Vatican City State, in Italian officially Stato della Città del Vaticano , which translates literally as State of the City of the Vatican, is a landlocked sovereign city-state whose territory consists of a walled enclave within the city of Rome, Italy. It has an area of...
- Northern Europe: Channel Islands
- Oceania
- Australasia: Jarrah ForestJarrah ForestJarrah Forest is an Interim Biogeographic Regionalisation for Australia region in Western Australia.-Location and description:The ecoregion stands on the 300m high Yilgarn block inland plateau and includes wooded valleys such as those of Western Australia's Murray River and the Helena River near...
, Swan Coastal PlainSwan Coastal PlainThe Swan Coastal Plain in Western Australia is the geographic feature which contains the Swan River as it travels west to the Indian Ocean. The coastal plain continues well beyond the boundaries of the Swan River and its tributaries, as a geological and biological zone, one of Western Australia's...
, WarrenWarren (biogeographic region)Warren, also known as Karri Forest Region and the Jarrah-Karri forest and shrublands ecoregion, is a biogeographic region in southern Western Australia. Located in the southwest corner of Western Australia between Cape Naturaliste and Albany, it is bordered to the north and east by the Jarrah...
, New ZealandNew ZealandNew Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...
- Australasia: Jarrah Forest
- Range maps for continents
Predators
The seed of Common groundsel is a good green foodGreen politics
Green politics is a political ideology that aims for the creation of an ecologically sustainable society rooted in environmentalism, social liberalism, and grassroots democracy...
for Canaries
Domestic Canary
The Domestic Canary, often simply known as the canary, is a domesticated form of the wild Canary, a small songbird in the finch family originating from the Macaronesian Islands ....
and Finches
British finches
The British finches are made up of several species of Finch which were formerly very popular as cage birds in Great Britain. Nowadays they are not commonplace, but are still kept by a few dedicated fanciers...
and it is available all year
Year
A year is the orbital period of the Earth moving around the Sun. For an observer on Earth, this corresponds to the period it takes the Sun to complete one course throughout the zodiac along the ecliptic....
round.
Senecio vulgaris seed has been found in the droppings of sparrow
Sparrow
The sparrows are a family of small passerine birds, Passeridae. They are also known as true sparrows, or Old World sparrows, names also used for a genus of the family, Passer...
s, and seedlings have been raised from the excreta of various birds. Seed has also been found in cow manure
Cow dung
Cow dung is the waste product of bovine animal species. These species include domestic cattle , bison , yak and water buffalo. Cow dung is the undigested residue of plant matter which has passed through the animal's gut. The resultant faecal matter is rich in minerals...
.
Some Lepidoptera
Lepidoptera
Lepidoptera is a large order of insects that includes moths and butterflies . It is one of the most widespread and widely recognizable insect orders in the world, encompassing moths and the three superfamilies of butterflies, skipper butterflies, and moth-butterflies...
species eat many of the Senecio genus; additional studies via electrophysiological recordings
Electrophysiology
Electrophysiology is the study of the electrical properties of biological cells and tissues. It involves measurements of voltage change or electric current on a wide variety of scales from single ion channel proteins to whole organs like the heart...
have shown that the taste sensilla
Sensilla
A sensillum is an insect sensory organ protruding from the cuticle, or sometime lying within or beneath it. Sensilla are divided in chemical, mechanical , thermal and visual....
of the Cinnabar moth
Cinnabar moth
The Cinnabar moth is a brightly coloured arctiid moth, found in Europe and western and central Asia. It has been introduced into New Zealand, Australia and North America to control poisonous ragwort, which its larvae feed on. The moth is named after the red mineral cinnabar because of the red...
larvae
Caterpillar
Caterpillars are the larval form of members of the order Lepidoptera . They are mostly herbivorous in food habit, although some species are insectivorous. Caterpillars are voracious feeders and many of them are considered to be pests in agriculture...
respond (get excited) specifically to the pyrrolizidine alkaloids, which all Senecio are known to contain.
Moths and caterpillars
- Cinnabar mothCinnabar mothThe Cinnabar moth is a brightly coloured arctiid moth, found in Europe and western and central Asia. It has been introduced into New Zealand, Australia and North America to control poisonous ragwort, which its larvae feed on. The moth is named after the red mineral cinnabar because of the red...
(Tyria jacobaeae) - Flame ShoulderFlame ShoulderThe Flame Shoulder is a moth of the family Noctuidae. It is distributed throughout Europe.The forewings of this species are reddish brown with a black streak interrupted by white stigmata and a creamy-yellow streak along the costa which gives the species its common name. The hindwings are pure...
(Ochropleura plecta) - Ragwort plume mothPlume mothThe Pterophoridae or plume moths are a family of Lepidoptera with unusually modified wings. Though they belong to the Apoditrysia like the larger moths and the butterflies, unlike these they are tiny and were formerly included among the assemblage called "Microlepidoptera".-Description and...
(Platyptilia isodactyla)
The Senecio genus also are host to other insects:
Beetles
- Ragwort flea beetleFlea beetleFlea beetles is a general name applied to the small, jumping beetles of the leaf beetle family . They make up the tribe Alticini, which is a part of the subfamily Galerucinae, though they were historically classified as a subfamily in their own right...
(Longitarsus jacobaeaeLongitarsus jacobaeaeLongitarsus jacobaeae is a species of flea beetle known as the tansy ragwort flea beetle. It is used as an agent of biological pest control against the noxious weed known as ragwort ....
) - Longitarsus gracilis (family Coleoptera species Chrysomelidae)
Flies
- Seed flies (DipteraDipteraDiptera , or true flies, is the order of insects possessing only a single pair of wings on the mesothorax; the metathorax bears a pair of drumstick like structures called the halteres, the remnants of the hind wings. It is a large order, containing an estimated 240,000 species, although under half...
: MuscoideaMuscoideaMuscoidea is a superfamily of flies in the subsection Calyptratae....
)- Ragwort seed fly (AnthomyiidaeAnthomyiidaeAnthomyiidae is a large and diverse family of Muscoidea flies. Name came from Greek "anthos" + "myia" . Some species are commonly called "root-maggots", as the larvae are found in the stems and roots of various plants...
, Botanophila seneciella)
- Ragwort seed fly (Anthomyiidae
- Gall flies (DipteraDipteraDiptera , or true flies, is the order of insects possessing only a single pair of wings on the mesothorax; the metathorax bears a pair of drumstick like structures called the halteres, the remnants of the hind wings. It is a large order, containing an estimated 240,000 species, although under half...
: TephritidaeTephritidaeTephritidae is one of two fly families referred to as "fruit flies", the other family being Drosophilidae. Tephritidae does not include the biological model organisms of the genus Drosophila , which is often called the "common fruit fly". There are nearly 5,000 described species of tephritid...
):- Ensina sonchi
- Sphenella marginata
- Trupanea stellata
- Trypeta zoe
and other insects that are not listed here.
The ragwort flea beetle and ragwort seed fly have been approved and released for Senecio control in California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...
, Australia and elsewhere.
Fungus
Most Senecio, including S. squalidus are susceptible to rust and other fungus
Pezizomycetes
Pezizomycetes are a class of fungi within the phylum Ascomycota.Pezizomycetes are apothecial fungi, meaning that their spore producing/releasing bodies are typically disk-like, bearing on their upper surfaces a layer of cylindrical spore producing cells called asci, from which the spores are...
and mildews
Powdery mildew
Powdery mildew is a fungal disease that affects a wide range of plants. Powdery mildew diseases are caused by many different species of fungi in the order Erysiphales. It is one of the easier diseases to spot, as its symptoms are quite distinctive. Infected plants display white powdery spots on the...
:
- Rust fungus Uredinales
- Coleosporium tussilaginisColeosporium tussilaginisColeosporium tussilaginis is species of rust fungus in the Coleosporiaceae family. It is a plant pathogen....
- stems and leaves (ColeosporiaceaeColeosporiaceaeThe Coleosporiaceae are a family of rust fungi in the order Pucciniales. The family contains 6 genera and 131 species....
) - Puccinia lagenophoraePucciniaPuccinia is a genus of fungi. All species in this genus are obligate plant pathogens and are known as rusts .Examples of Puccinia rusts and the diseases they cause:* Puccinia asparagi - Asparagus rust...
- leaves only (PucciniaceaePucciniaceaeThe Pucciniaceae are a family of rust fungi that cause plant diseases, mainly on cereals such as wheat. The family contains 20 genera and over 4900 species....
) - Bremia lactucaeBremia lactucaeBremia lactucae is a plant pathogen.- External links :* *...
- Coleosporium tussilaginis
- White rust PeronosporalesPeronosporalesThe Peronosporales are an order of water moulds which can be pathogenic.Many diseases of plants are sometimes classified under this order, but are sometimes considered members of order Pythiales. Some of these pathogenic protists include the organisms responsible for potato blight, eucalyptus...
- Albugo tragopogonisAlbugo tragopogonisAlbugo tragopogonis, Sunflower White Rust, is an oomycete plant pathogen, although some sources still regard it as a fungal organism.Albugo tragopogonis is the old name for Pustula tragopogonis, the causal agent of white blister disease on goatsbeard...
- (Albuginaceae) - some of the species PeronosporaceaePeronosporaceaePeronosporaceae are a family of water moulds that contains 17 genera, comprising more than 600 species. Most of them are called downy mildews. The closest relative of Peronosporaceae is Phytophthora....
- (Albuginaceae)
- Albugo tragopogonis
- Sac fungus AscochytaAscochytaAscochyta is a genus of ascomycote fungi, containing several pathogens to plants. The taxonomy of this genus is still incomplete.Some of these pathogens affect grass species, including grains.- Some species of Ascochyta :...
, PezizomycetesPezizomycetesPezizomycetes are a class of fungi within the phylum Ascomycota.Pezizomycetes are apothecial fungi, meaning that their spore producing/releasing bodies are typically disk-like, bearing on their upper surfaces a layer of cylindrical spore producing cells called asci, from which the spores are...
- Ascochyta senecionicola - (Coelomycete)
- Groundsel Mildew ErysiphalesErysiphalesErysiphales are an order of ascomycete fungi. The order contains one family, Erysiphaceae. Many of them cause plant diseases called powdery mildew.-Systematics:The order contains one family , 28 genera and approximately 100 species...
- Golovinomyces cichoracearum var. fischeri
- Powdery Mildew ErysiphalesErysiphalesErysiphales are an order of ascomycete fungi. The order contains one family, Erysiphaceae. Many of them cause plant diseases called powdery mildew.-Systematics:The order contains one family , 28 genera and approximately 100 species...
- Podosphaera fuscaPodosphaeraPodosphaera is a genus of fungi in the family Erysiphaceae. Species in this genus are plant pathogens, causing powdery mildew.-Species:*P. aphanis*P. balsaminae*P. clandestina var. aucupariae*P. clandestina var. clandestina...
- (ErysiphaceaeErysiphaceaeThe Erysiphaceae are fungal family of the order Erysiphales. The family contains many of the powdery mildews.-Genera:*Arthrocladiella*Blumeria*Brasiliomyces*Bulbomicrosphaera*Bulbouncinula*Caespitotheca...
)
- Podosphaera fusca
- Black root rot MicroascalesMicroascalesThe Microascales are an order of fungi in the Sordariomycetes class, subclass Hypocreomycetidae. This is a relatively small order of mostly saprobic fungi that live in soil, rotting vegetation and dung. Some species are plant pathogens, such as Ceratocystis fimbriata, transmitted by beetles to...
- some of the family Incertae sedisIncertae sedis, is a term used to define a taxonomic group where its broader relationships are unknown or undefined. Uncertainty at specific taxonomic levels is attributed by , , and similar terms.-Examples:*The fossil plant Paradinandra suecica could not be assigned to any...
- some of the family Incertae sedis
and other fungus that are not listed here.
Reputation for being noxious and toxic
Senecio vulgaris has been listed as a noxious weed,being both non-indigenous
Introduced species
An introduced species — or neozoon, alien, exotic, non-indigenous, or non-native species, or simply an introduction, is a species living outside its indigenous or native distributional range, and has arrived in an ecosystem or plant community by human activity, either deliberate or accidental...
to most if not all of the Americas and having a reputation for being hepatotoxic
Hepatotoxicity
Hepatotoxicity implies chemical-driven liver damage.The liver plays a central role in transforming and clearing chemicals and is susceptible to the toxicity from these agents. Certain medicinal agents, when taken in overdoses and sometimes even when introduced within therapeutic ranges, may injure...
to livestock
Livestock
Livestock refers to one or more domesticated animals raised in an agricultural setting to produce commodities such as food, fiber and labor. The term "livestock" as used in this article does not include poultry or farmed fish; however the inclusion of these, especially poultry, within the meaning...
and to humans.
Toxic versus medicinal
HumanAs a plant that is reported to be both poison
Poison
In the context of biology, poisons are substances that can cause disturbances to organisms, usually by chemical reaction or other activity on the molecular scale, when a sufficient quantity is absorbed by an organism....
ous for human ingestion
Digestion
Digestion is the mechanical and chemical breakdown of food into smaller components that are more easily absorbed into a blood stream, for instance. Digestion is a form of catabolism: a breakdown of large food molecules to smaller ones....
and also medicinal; much of the contradiction can be found by closely reviewing the words that are used and the dose (amount) of the poisonous substance that is ingested to prove either claim. All species of the genus Senecio contain pyrrolizidine alkaloids
Pyrrolizidine
Pyrrolizidine alkaloids are a group of naturally occurring alkaloids based on the structure of pyrrolizidine. Pyrrolizidine alkaloids are produced by plants as a defense mechanism against insect herbivores. More than 660 PAs and PA N-oxides have been identified in over 6,000 plants, and about...
(e.g., senecionine
Senecionine N-oxygenase
In enzymology, a senecionine N-oxygenase is an enzyme that catalyzes the chemical reactionThe 4 substrates of this enzyme are senecionine, NADPH, H+, and O2, whereas its 3 products are senecionine N-oxide, NADP+, and H2O....
) a substance that when a human has chronic exposure
can cause irreversible liver damage.
Common groundsel as a medicinal herb does not seem to be recommended very often since 1931, when it was recommended as a diaphoretic
Diaphoresis
Diaphoresis is excessive sweating commonly associated with shock and other medical emergency conditions.Diaphoretic is the state of perspiring profusely, or something that has the power to cause increased perspiration....
, an antiscorbutic
Scurvy
Scurvy is a disease resulting from a deficiency of vitamin C, which is required for the synthesis of collagen in humans. The chemical name for vitamin C, ascorbic acid, is derived from the Latin name of scurvy, scorbutus, which also provides the adjective scorbutic...
, a purgative
Laxative
Laxatives are foods, compounds, or drugs taken to induce bowel movements or to loosen the stool, most often taken to treat constipation. Certain stimulant, lubricant, and saline laxatives are used to evacuate the colon for rectal and/or bowel examinations, and may be supplemented by enemas under...
, a diuretic
Diuretic
A diuretic provides a means of forced diuresis which elevates the rate of urination. There are several categories of diuretics. All diuretics increase the excretion of water from bodies, although each class does so in a distinct way.- Medical uses :...
and an anthelmintic
Anthelmintic
Anthelmintics or antihelminthics are drugs that expel parasitic worms from the body, by either stunning or killing them. They may also be called vermifuges or vermicides .-Pharmaceutical classes:...
, which was a demotion as it was previously suggested for the expelling of gravel of the kidneys and reins
Kidney stone
A kidney stone, also known as a renal calculus is a solid concretion or crystal aggregation formed in the kidneys from dietary minerals in the urine...
by Pedanius Dioscorides
Pedanius Dioscorides
Pedanius Dioscorides was a Greek physician, pharmacologist and botanist, the author of a 5-volume encyclopedia about herbal medicine and related medicinal substances , that was widely read for more than 1,500 years.-Life:...
in the 70s-90s, for use as poultice
Poultice
A poultice, also called cataplasm, is a soft moist mass, often heated and medicated, that is spread on cloth over the skin to treat an aching, inflamed, or painful part of the body. It can be used on wounds such as cuts...
s by John Gerard
John Gerard
John Gerard aka John Gerarde was an English herbalist notable for his herbal garden and botany writing. In 1597 he published a large and heavily illustrated "Herball, or Generall Historie of Plantes", which went on to be the most widely circulated botany book in English in the 17th century...
in the late 16th century and as a cure for epilepsy
Epilepsy
Epilepsy is a common chronic neurological disorder characterized by seizures. These seizures are transient signs and/or symptoms of abnormal, excessive or hypersynchronous neuronal activity in the brain.About 50 million people worldwide have epilepsy, and nearly two out of every three new cases...
by Nicholas Culpeper
Nicholas Culpeper
Nicholas Culpeper was an English botanist, herbalist, physician, and astrologer. His published books include The English Physician and the Complete Herbal , which contain a rich store of pharmaceutical and herbal knowledge, and Astrological Judgement of Diseases from the Decumbiture of the Sick ,...
in the 17th century. More current information is contradictory about the dangers of the ingestion of Groundsel. A heavily referenced paper from 1989 suggests that the response is immediate and gives pre-ambulatory care recommendations. A Canadian poisonous plants information database references a paper from 1990 in presenting this prenatal warning: "In a case of prenatal exposure, a mother ingested tea containing an estimated 0.343 milligram of senecionine, resulting in fatal veno-occlusive disease in a newborn infant." Information about the pyrrolizidine alkaloids, the substance present in Senecio vulgaris is much less contradictory and all warn of accumulation of the alkaloid.
Certain pyrrolizidine alkaloids are non-toxic precursors that are converted to toxic metabolite
Metabolite
Metabolites are the intermediates and products of metabolism. The term metabolite is usually restricted to small molecules. A primary metabolite is directly involved in normal growth, development, and reproduction. Alcohol is an example of a primary metabolite produced in large-scale by industrial...
s in the body in a process called toxification
Toxification
Toxification is the conversion of chemical compounds into more toxic forms. This conversion can take place in the body as well as in substrates such as soil or water...
Livestock
Linnaeus
Carolus Linnaeus
Carl Linnaeus , also known after his ennoblement as , was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology...
is cited to have claimed that "goat
Goat
The domestic goat is a subspecies of goat domesticated from the wild goat of southwest Asia and Eastern Europe. The goat is a member of the Bovidae family and is closely related to the sheep as both are in the goat-antelope subfamily Caprinae. There are over three hundred distinct breeds of...
s and swine
Pig
A pig is any of the animals in the genus Sus, within the Suidae family of even-toed ungulates. Pigs include the domestic pig, its ancestor the wild boar, and several other wild relatives...
eat this common plant freely, cows
Cattle
Cattle are the most common type of large domesticated ungulates. They are a prominent modern member of the subfamily Bovinae, are the most widespread species of the genus Bos, and are most commonly classified collectively as Bos primigenius...
being not partial to it and horse
Horse
The horse is one of two extant subspecies of Equus ferus, or the wild horse. It is a single-hooved mammal belonging to the taxonomic family Equidae. The horse has evolved over the past 45 to 55 million years from a small multi-toed creature into the large, single-toed animal of today...
s and sheep
Domestic sheep
Sheep are quadrupedal, ruminant mammals typically kept as livestock. Like all ruminants, sheep are members of the order Artiodactyla, the even-toed ungulates. Although the name "sheep" applies to many species in the genus Ovis, in everyday usage it almost always refers to Ovis aries...
declining to touch it, but not only are caged birds
Birdcage
A birdcage is a cage designed to house birds as pets.Antique birdcages are often popular as collectors' items or as household decor but most are not suitable for housing live birds, being too small, or of unsafe materials or construction...
fond of it (the seeds), but its leaves and seeds afford food for many of our wild species (rabbit
Rabbit
Rabbits are small mammals in the family Leporidae of the order Lagomorpha, found in several parts of the world...
s were given as an example)." More recent studies claim that the lethal
Lethal
Lethal is an adjective meaning capable of causing death.Lethal may also refer to:* Lethal , an American heavy metal band* Leigh Matthews, Australian rules footballer nicknamed "Lethal" or "Lethal" Leigh...
amount that cattle or horses need to consume is 7% of their body weight
Body weight
The term body weight is used in daily English speech as well as in the contexts of biological and medical sciences to describe the mass of an organism's body. Body weight is measured in kilograms throughout the world, although in some countries it is still measured in pounds or stones and pounds...
(example: 50 pounds (23 kg) would need to be consumed by a cow weighing 700 pounds (318 kg)). Lesser amounts cause the liver
Liver
The liver is a vital organ present in vertebrates and some other animals. It has a wide range of functions, including detoxification, protein synthesis, and production of biochemicals necessary for digestion...
to lose function but is not apparent until the animal is stressed (by new feed or location, pregnancy, a different toxin
Toxin
A toxin is a poisonous substance produced within living cells or organisms; man-made substances created by artificial processes are thus excluded...
, etc.). Sheep and goats have rumen
Rumen
The rumen, also known as a paunch, forms the larger part of the reticulorumen, which is the first chamber in the alimentary canal of ruminant animals. It serves as the primary site for microbial fermentation of ingested feed...
bacteria
Bacteria
Bacteria are a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria have a wide range of shapes, ranging from spheres to rods and spirals...
that detoxify the alkaloid
Alkaloid
Alkaloids are a group of naturally occurring chemical compounds that contain mostly basic nitrogen atoms. This group also includes some related compounds with neutral and even weakly acidic properties. Also some synthetic compounds of similar structure are attributed to alkaloids...
s and are able to consume twice their body weight of this and other species of genus Senecio. The alkaloids responsible are not destroyed by drying or by fermentation in silage.
Introduced versus invasive
Introduced species become invasive when they compete with nativesIndigenous (ecology)
In biogeography, a species is defined as native to a given region or ecosystem if its presence in that region is the result of only natural processes, with no human intervention. Every natural organism has its own natural range of distribution in which it is regarded as native...
or with crops
Crop (agriculture)
A crop is a non-animal species or variety that is grown to be harvested as food, livestock fodder, fuel or for any other economic purpose. Major world crops include maize , wheat, rice, soybeans, hay, potatoes and cotton. While the term "crop" most commonly refers to plants, it can also include...
. Senecio vulgaris is not known to be a strong competitor but it has been known to reduce mint production.
There is evidence that it is not a strong invasive and sometimes protective of critically endangered native plants.
Control
The approximately 22 millimetre (0.866141732283465 in) long
pappus
Pappus (flower structure)
The pappus is the modified calyx, the part of an individual disk, ray or ligule floret surrounding the base of the corolla, in flower heads of the plant family Asteraceae. The pappus may be composed of bristles , awns, scales, or may be absent. In some species, the pappus is too small to see...
seeds of Senecio vulgaris, each plant capable of producing 25,000 or more seeds (1,700 seeds per plant are more likely) with three generations of the plant per year; seeds that are widely dispersed by the wind,
have been identified as a contaminant of cereal and vegetable seeds and a poison to some livestock; there is some inspiration to understand the growth stages and determine some control methods.
- Cultivation: Cultivation with the handHoe (tool)A hoe is an ancient and versatile agricultural tool used to move small amounts of soil. Common goals include weed control by agitating the surface of the soil around plants, piling soil around the base of plants , creating narrow furrows and shallow trenches for planting seeds and bulbs, to chop...
or tiller is a recommended method of controlling Senecio vulgaris from growing in gardens and planting fields; cultivate to a depth of 2 inches (50.8 mm). The plant does prefer to take root in disturbed soils, so cultivation rids new plants but also buries and stirs up new seeds so the cultivation needs to be repeated at 14-day intervals. Seeds can still mature even when the plant has been killed; seed from plants cut in flower had germination levels of 35%. Groundsel seed numbers increased in soil during a 2-year set-aside left fallow but not when there was a sown grass cover. The weed cannot live on grazed, trampled or mowed sites.
- Biological: The pathogenPathogenA pathogen gignomai "I give birth to") or infectious agent — colloquially, a germ — is a microbe or microorganism such as a virus, bacterium, prion, or fungus that causes disease in its animal or plant host...
rust fungusRust (fungus)Rusts are plant diseases caused by pathogenic fungi of the order Pucciniales. About 7800 species are known. Rusts can affect a variety of plants; leaves, stems, fruits and seeds. Rust is most commonly seen as coloured powder, composed off tiny aeciospores which land on vegetation producing...
or Puccinia lagenophoraePucciniaceaeThe Pucciniaceae are a family of rust fungi that cause plant diseases, mainly on cereals such as wheat. The family contains 20 genera and over 4900 species....
and the Cinnabar mothCinnabar mothThe Cinnabar moth is a brightly coloured arctiid moth, found in Europe and western and central Asia. It has been introduced into New Zealand, Australia and North America to control poisonous ragwort, which its larvae feed on. The moth is named after the red mineral cinnabar because of the red...
Tyria jacobaeae have both been used and studied in an attempt to control infestation of Senecio vulgaris. One study showed that rust fungus infected Senecio vulgaris survived and actually used more of the available soil nutrients. The cinnabar moth eats groundsel between June and August, but the seeds germinate and the plant grows as soon as the ground is warm enough (and after a warm rain), making this an insufficient control almost everywhere groundsel can be found.
- Chemical: Herbicides designed to control broadleaf plantsFlowering plantThe flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...
are affective for controlling Senecio vulgaris in cerealCerealCereals are grasses cultivated for the edible components of their grain , composed of the endosperm, germ, and bran...
s and forage grassesFodderFodder or animal feed is any agricultural foodstuff used specifically to feed domesticated livestock such as cattle, goats, sheep, horses, chickens and pigs. Most animal feed is from plants but some is of animal origin...
but also will "control" broadleaf cropsVegetableThe noun vegetable usually means an edible plant or part of a plant other than a sweet fruit or seed. This typically means the leaf, stem, or root of a plant....
, such as mintMenthaMentha is a genus of flowering plants in the family Lamiaceae . The species are not clearly distinct and estimates of the number of species varies from 13 to 18. Hybridization between some of the species occurs naturally...
, forage legumes, strawberriesStrawberryFragaria is a genus of flowering plants in the rose family, Rosaceae, commonly known as strawberries for their edible fruits. Although it is commonly thought that strawberries get their name from straw being used as a mulch in cultivating the plants, the etymology of the word is uncertain. There...
, carrotCarrotThe carrot is a root vegetable, usually orange in colour, though purple, red, white, and yellow varieties exist. It has a crisp texture when fresh...
s and all other non-grass crops. There is also evidence that the plant develops an immunity to the chemical controlMutagenIn genetics, a mutagen is a physical or chemical agent that changes the genetic material, usually DNA, of an organism and thus increases the frequency of mutations above the natural background level. As many mutations cause cancer, mutagens are therefore also likely to be carcinogens...
.
- Other: Groundsel seedlings with 2-6 leaves are tolerant of flame weeding but the seeds are susceptible to soil solarizationSoil solarizationSoil solarization is an environmentally friendly method of using solar power for controlling disease agents in the soil by mulching the soil and covering it with tarp, usually with a transparent polyethylene cover, to trap solar energy....
.
Subspecies synonyms
- Senecio vulgaris subsp. denticulatus (O.F. Muell.) P.D. Sell
- Senecio vulgaris var. dubius (Ledeb.) Franch., 1883
- Senecio vulgaris var. dubius Trautv., 1866
- Senecio vulgaris var. hibernicus Syme
- Senecio vulgaris var. vulgaris
- Senecio vulgaris-humilis Batt. & Trab.